Evieux Formation

The Evieux Formation, formerly included within the “Bois-de-Mouches Formation” and containing the Strud locality, is a late Devonian formation from Belgium. It preserves a wide range of fossils, including some of the earliest branchiopod crustaceans, various fish, and the putative insect Strudiella.^[1]

Evieux Formation
Stratigraphic range: Famennian
Type	Geological formation
Underlies	Dolhain Formation [nl]/Comblain-au-Pont Formation[1]
Overlies	Monfort Formation/Citadelle de Huy Formation[1]
Location
Region	Namur Province
Country	Belgium

Paleobiota

Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon		Notes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.

Paleobiota
Genus	Species	Higher taxon	Notes	Images
Strudiella	S. devonica	Mandibulata	Debated whether it is an insect or a non-insect crustacean.[2][3]	Interpretation of Strudiella
Strudops	S. goldenbergi	Notostraca	The earliest notostracan known[4][5]	Strudops reconstruction
Haltinnaias	H. serrata	Anostraca	The earliest fairy shrimp known[5]
Gesvesia	G. pernegrei	Spinicaudata	One of the few diplostracan fossils with preserved internal anatomy[5]
Tealliocaris	T. walloniensis	Pygocephalomorpha	The earliest freshwater decapod known[6]	T. etheridgii reconstruction
Schramidontus	S. labasensis	Angustidontida	One of only two angustidontids, alongside the type genus Angustidontus.[7]	Reconstruction of Angustidontus
Nervicaris	N. belgica[8]	Echinocarididae (Phyllocarida)
Hardieopteridae indet.	Unapplicable	Kokomopteroidea	Several juvenile specimens, not assignable to any genus[9]
Ericixerxes	E. pottii	Euthycarcinoidea	Likely breathed air as evidenced by unusual circular structures in the fossil’s abdomen[10]
Microconchus	M. helicteres	Microconchida	Lived in lagoonal ponds[11]	Microconchus from the Devonian of Ohio
Grossilepis [species]	G. rikiki	Bothriolepididae	Also known from a locality near Moresnet[12]
Turrisaspis	T. strudensis	Groenlandaspididae	Small size of specimens shows Strud was likely a placoderm “nursery”[13]	T. elektor reconstruction
Groenlandaspis	G. potyi[14]	Groenlandaspididae	One of the few known Paleozoic genera with preserved colour	G. antarctica illustration
Phyllolepis	P. undulata	Phyllolepididae	Synonymised with several other species[15]	Reconstruction of Phyllolepis
Remigolepis	R. ?durnalensis	Antiarchi	Known from the Becco locality, as opposed to Strud[14]	R. zhongweiensis reconstruction
Diplacanthidae indet.	Unapplicable	Acanthodii	Known from two large spines[14]
Holoptychius[1]	H. flemingi	Porolepiformes		Life restoration of Holoptychius sp.
Soederberghia	S. sp[14]	Dipnoi	A relatively widespread lungfish[16]
Jarvikia?	?J. sp[16]	Dipnoi
Langlieria	L. socqueti[17]	Tristichopteridae	Also known from the Catskill Formation	Jawbone of Langlieria
Eusthenodon	E. wangsjoi[18]	Tristichopteridae	Known from all over the world	Life restoration of E. wangsjoi
Stegocephalia indet.	Unapplicable	Elpistostegalia	Fragmentary remains, with Ichthyostega-like jawbones and a whatcheeriid-like postorbital bone[19]
Moresnetia	M. zalesskyi[14]	Spermatophyta	One of the earliest seed plants[20]
Dorinnotheca	D. streelii	Lyginopteridales	One of the earliest known seed plants[21]
Condrusia	C. rumex	Spermatophyta[14]	Gained a lectotype in 2006[22]	C. rumex fossil
Thorezia	T. verezensis[23]	Spermatophyta
Archaeopteris	A. roemeriana[24]	Progymnospermopsida	One of the most widespread Devonian trees	Fossil of A. hibernica
Barinophyton	B. citrulliforme[25]	Barinophytes	Known from various other localities	B. citrulliforme fossil from Pennsylvania
Aneurophyton	A. olnense[25]	Aneurophytales	Basal progymnosperm	Fossil of A. sp
Rhacophyton	R. condrusorum	Rhacophytales	One of the most abundant plants at Evieux[25]	Rhacophyton sp fossil
Eviostachya	E. sp[25]	Sphenophyllales
Sphenophyllum	S. subtenerrimum[25]	Sphenophyllales
Barsostrobus	B. famennensis[25][26]	Lycophyta