Densuș-Ciula Formation

The Densuș-Ciula Formation is a geological formation in Romania whose strata date back to the Late Cretaceous. It forms part of the Hațeg Island assemblage. Dinosaur remains are among the fossils that have been recovered from the formation.^[1] It is divided up into three members, the lower member is noted for high content of volcanogenic material and is poorly fossiliferous. While the Middle member consists of silty mudstones, sandstones and conglomerates containing volcanogenic clasts and is richly fossiliferous, while the upper member consists of matrix supported red conglomerates and is poorly fossiliferous.^[2]

Densuș-Ciula Formation
Stratigraphic range: Maastrichtian PreꞒ Ꞓ O S D C P T J K Pg N
Type	Geological formation
Sub-units	Lower, Middle and Upper Members
Underlies	Cenozoic deposits
Overlies	Flysch
Lithology
Primary	Mudstone, sandstone, conglomerate
Location
Country	Romania
Extent	Northwest Hațeg Basin

Fossil content

Indeterminate dromaeosaurid and possible indeterminate troodontid remains present in Județul Hunedoara, Romania.^[1] An unnamed theropod is also present.^[1]

Amphibians

Amphibians
Genus	Species	Presence	Notes	Images
Albanerpeton	Indeterminate		Albanerpetontid
Paralatonia	P. transylvanica		Alytid
Hatzegobatrachus	H. grigorescui		Bombinatorid

Squamates

Squamates
Genus	Species	Presence	Notes	Images
Nidophis	N. insularis		Madtsoiid snake

Turtles

Turtles
Genus	Species	Presence	Notes	Images
Kallokibotion	K. bajazidi
Dortokidae	Indeterminate

Crocodyliformes

Crocodyliformes
Genus	Species	Presence	Notes	Images
Allodaposuchus	A. precedens
Doratodon	Indeterminate
Sabresuchus	S. sympiestodon		Formerly thought to be Theriosuchus

Dinosaurs

Ornithischians

Ornithischians of the Densuș-Ciula Formation
Genus	Species	Presence	Material	Notes	Images
Telmatosaurus	T. transsylvanicus		A skull.	A hadrosauromorph.
Zalmoxes	Z. robustus	Hunedoara County		A small rhabdodontid ornithopod[1]
	Z. shqiperorum	Hunedoara County

Sauropods

Sauropods of the Densuș-Ciula Formation
Genus	Species	Presence	Material	Notes	Images
Uriash	U. kadici[3]	Locality VI of Kadić.	A partial skeleton composed of incomplete vertebrae, metatarsal, femur, and humerus.	A titanosaur.

Theropods

Theropods of theDensuș-Ciula Formation
Genus	Species	Presence	Material	Notes	Images
Balaur	B. bondoc			A paravian probably a junior synonym of Elopteryx[4][5]

Pterosaurs

Pterosaurs of Densuș-Ciula Formation.
Genus	Species	Presence	Material	Notes	Images
Hatzegopteryx[6]	H. thambema[6]	Valioara	A preserved robust skull, incomplete left humerus, and unidentified bone fragments.	A giant azhdarchid pterosaur.

Mammals

Mammals
Genus	Species	Presence	Notes	Images
Hainina	Indeterminate		Two species

See also

• List of dinosaur-bearing rock formations