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Nipponopterus
Genus of azhdarchid pterosaur from the Late Cretaceous From Wikipedia, the free encyclopedia
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Nipponopterus (meaning "Nippon wing") is an extinct genus of azhdarchid pterosaur that lived during the Late Cretaceous (Turonian to Coniacian ages) in what is now Japan. It is known from a partial cervical (neck) vertebra found in the Mifune Group, located in the Kumamoto Prefecture in Kyūshū. The genus contains a single species, Nipponopterus mifunensis, named and described in 2025. It is the first pterosaur to be named from Japan.
Morphologically, Nipponopterus is similar to a Mongolian specimen nicknamed the "Burkhant azhdarchid". Both are estimated to have had a wingspan measuring around 3 to 3.5 m (9 ft 10 in to 11 ft 6 in) when fully grown. Nipponopterus is a member of the clade Quetzalcoatlinae within the family Azhdarchidae.
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Discovery and naming
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Nipponopterus type locality in the Mifune Group of Japan.
The holotype specimen of Nipponopterus, MDM 349, was discovered in sediments of the "Upper Formation" of the Mifune Group ('Locality 1018') in rock outcrops near Amagimi Dam in Mifune Town, Kumamoto Prefecture in Kyūshū, Japan. The specimen consists of the posterior end of the sixth cervical (neck) vertebra. The specimen was first described in 2000 as belonging to an indeterminate member of the family Azhdarchidae. The limited understanding of the clade at that time precluded more detailed analyses.[1] While early reviews interpreted the cervical vertebra as the fourth[2] or fifth,[3] the most recent analysis suggests it can confidently be regarded as the sixth.[4]
In 2025, Chinese paleontologist Xuanyu Zhou and colleagues named and described Nipponopterus mifunensis as a new genus and species of azhdarchid pterosaur based on these fossil remains. The generic name Nipponopterus combines "Nippon"—the Japanese name for the country—with "pterus", derived from the Ancient Greek word πτερόν (ptéron), meaning "wing". The specific name mifunensis refers to the type locality in the Mifune Group in Mifune Town. Nipponopterus is the first pterosaur named from Japan.[4]
A wing phalanx and metacarpal belonging to indeterminate azhdarchid pterosaurs are also known from this formation, but have not been referred to Nipponopterus.[4]
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Description
The holotype specimen of Nipponopterus is morphologically comparable to a similarly fragmentary unnamed pterosaur specimen from Mongolia, nicknamed the "Burkhant azhdarchid".[5] Measurements of the material of Nipponopterus indicates that it is 82% the size of the Burkhant specimen. However, the former likely belongs to a subadult individual, meaning it was not fully grown. As such, Zhou and colleagues, in their 2025 description of Nipponopterus, estimated that both pterosaurs would have had similar wingspans, at around 3 to 3.5 m (9 ft 10 in to 11 ft 6 in).[4]
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Classification
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In their phylogenetic analysis using the dataset of paleontologist Rodrigo Pêgas in 2024,[6] Zhou and colleagues recovered Nipponopterus as a member of the clade Quetzalcoatlinae within Azhdarchidae. It was placed as the sister taxon to the "Burkhant azhdarchid", which corroborates their close relationship in terms of similar features and geography.[4] The results of their study are displayed in the cladogram below:
| Azhdarchidae |
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References
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