Phalangiotarbida

Phalangiotarbida is an extinct arachnid order first recorded from the Early Devonian of Germany and most widespread in the Upper Carboniferous coal measures of Europe and North America.^[1] The last species are known from the early Permian Rotliegend of Germany.

Phalangiotarbida Temporal range: Early Devonian–Early Permian PreꞒ Ꞓ O S D C P T J K Pg N
Goniotarbus angulatus fossil, dorsal and ventral views
Reconstruction of Goniotarbus angulatus and Mesotarbus peteri
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Subphylum:	Chelicerata
Class:	Arachnida
Order:	†Phalangiotarbida Haase, 1890
Families
Currently 4 families, see text for details.
Synonyms
Architarbi Phalangiotarbi

The affinities of phalangiotarbids are obscure, with most authors favouring affinities with Opiliones (harvestmen) and/or Acari (mites and ticks). Phalangiotarbida has been recently (2004) proposed to be sister group to (Palpigradi+Tetrapulmonata): the taxon Megoperculata sensu Shultz (1990).^[2]

Nemastomoides depressus, described as a harvestman in the family Nemastomoididae, is actually a poorly preserved phalangiotarbid.^[3]

Taxa included

• Family Anthracotarbidae Kjellesvig-Waering, 1969
  • Genus Anthracotarbus Kjellesvig-Waering, 1969
    • Species Anthracotarbus hintoni Kjellesvig-Waering, 1969
• Family Architarbidae Karsch, 1882
  • Genus Architarbus Scudder, 1868
    • Species Architarbus hoffmanni Guthörl, 1934 (Jr synonyms Opiliotarbus kliveri Waterlot, 1934; Goniotarbus sarana Guthörl, 1965)
    • Species Architarbus minor Petrunkevitch, 1913
    • Species Architarbus rotundatus Scudder, 1868
  • Genus Bornatarbus Rößler & Schneider, 1997
    • Species Bornatarbus mayasii Haupt in Nindel, 1955
  • Genus Devonotarbus Poschmann, Anderson & Dunlop, 2005
    • Species Devonotarbus hombachensis Poschmann, Anderson & Dunlop, 2005
  • Genus Discotarbus Petrunkevitch, 1913
    • Species Discotarbus deplanatus Petrunkevitch, 1913
  • Genus Geratarbus Scudder, 1890
    • Species Geratarbus lacoei Scudder, 1890
    • Species Geratarbus bohemicus Petrunkevitch, 1953
  • Genus Goniotarbus Petrunkevitch, 1953
    • Species Goniotarbus angulatus Pocock, 1911
    • Species Goniotarbus tuberculatus Pocock, 1911
  • Genus Hadrachne Melander, 1903
    • Species Hadrachne horribilis Melander, 1903
  • Genus Leptotarbus Petrunkevitch, 1945
    • Species Leptotarbus torpedo Pocock, 1911
  • Genus Mesotarbus Petrunkevitch, 1949
    • Species Mesotarbus angustus Pocock, 1911
    • Species Mesotarbus eggintoni Pocock, 1911
    • Species Mesotarbus hindi Pocock, 1911
    • Species Mesotarbus intermedius Petrunkevitch, 1949
    • Species Mesotarbus peteri Dunlop & Horrocks, 1997
  • Genus Metatarbus Petrunkevitch, 1913
    • Species Metatarbus triangularus Petrunkevitch, 1913
  • Genus Ootarbus Petrunkevitch, 1945
    • Species Ootarbus pulcher Petrunkevitch, 1945
    • Species Ootarbus ovatus Petrunkevitch, 1945
  • Genus OrthotarbusPetrunkevitch, 1945
    • Species Orthotarbus minutus Petrunkevitch, 1913
    • Species Orthotarbus robustus Petrunkevitch, 1945
    • Species Orthotarbus nyranensis Petrunkevitch, 1953
  • Genus Paratarbus Petrunkevitch, 1945
    • Species Paratarbus carbonarius Petrunkevitch, 1945
  • Genus Phalangiotarbus Haase, 1890
    • Species Phalangiotarbus subovalis Woodward, 1872
  • Genus Pycnotarbus Darber, 1990
    • Species Pycnotarbus verrucosus Darber, 1990
  • Genus Triangulotarbus Patrick, 1989
    • Species Triangulotarbus terrehautensis Patrick, 1989
• Family Heterotarbidae Petrunkevitch, 1913
  • Genus Heterotarbus Petrunkevitch, 1913
    • Species Heterotarbus ovatus Petrunkevitch, 1913
• Family Opiliotarbidae Petrunkevitch, 1949
  • Genus Opiliotarbus Pocock, 1910
    • Species Opiliotarbus elongatus Scudder, 1890
• nomina dubia
  • Eotarbus litoralis Kušta, 1888
  • Nemastomoides depressus Petrunkevitch, 1913