Portugalosuchus

Portugalosuchus (meaning "crocodile from Portugal") is an extinct genus of eusuchian crocodyliform that was possibly a basal crocodylian – if so then it would be the oldest known crocodylian to date.^[2] The type species is P. azenhae, described in 2018,^[2] and it is known from the Late Cretaceous (Cenomanian)-aged Tentugal Formation in Portugal.^[3] A 2021 morphological study recovered Portugalosuchus within Crocodylia as a member of Gavialidae closely related to similar "thoracosaurs" (e.g. Thoracosaurus), while also noting that it might also possibly be outside of Crocodylia completely.^[1] A 2022 tip dating analysis incorporating both morphological and DNA data placed Portugalosuchus outside of Crocodylia, as the sister taxon of the family Allodaposuchidae. A cladogram simplified after that analysis is shown below:^[4]

Eusuchia	Hylaeochampsa Acynodon Iharkutosuchus Portugalosuchus Agaresuchus Lohuecosuchus Allodaposuchus Arenysuchus "Thoracosaurs" Borealosuchus Planocraniidae Crocodylia

Portugalosuchus Temporal range: Late Cretaceous: Cenomanian, 99.7–94.3 Ma[1] PreꞒ Ꞓ O S D C P T J K Pg N
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Clade:	Archosauria
Clade:	Pseudosuchia
Clade:	Crocodylomorpha
Genus:	†Portugalosuchus Mateus et al., 2018
Species:	†P. azenhae
Binomial name
†Portugalosuchus azenhae Mateus et al., 2018

The morphology-based phylogenetic analyses based on the new neuroanatomical data obtained from its skull using micro-CT scans suggested that this taxon is a crown group crocodilian and a member of the 'thoracosaurs', recovered as a sister taxon of Thoracosaurus within Gavialoidea,^[5] though it is uncertain whether 'thoracosaurs' were true gavialoids.^[6]