Prodidomidae

Prodidomidae is a family of spider, sometimes called long-spinneret ground spiders. It was formerly regarded as a subfamily of Gnaphosidae, but was raised to a family in 2022.^[1]

Long-spinneret ground spiders
Zimiris doriae
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Subphylum:	Chelicerata
Class:	Arachnida
Order:	Araneae
Infraorder:	Araneomorphae
Family:	Prodidomidae Simon, 1884
Genera
See text.

Spiders in the family are easily identified by the greatly elongated base of the piriform gland spigots. At least parts of their body are covered with shiny scales or setae. The posterior median eyes are flat and silvery, with a triangular, egg-shaped or irregularly rectangular shape.^[2]

Biology

Spiders in the Prodidomidae are ground dwellers. Most species are nocturnal and hide during the day in litter, but Myandra species, which are probably mimicking ants, seem to be active during the day.^[2] The genus Zimiris is synanthropic and thus found throughout the tropics.

Distribution

Although Theuma walteri was described from Turkmenistan by Eugène Simon, it is suspected that Simon accidentally exchanged its locality with that of Anagraphis pallens (Gnaphosidae); then T. walteri would have been collected in the Cape of Good Hope, while A. pallens is from Turkmenistan.^[3]

Genera

As of September 2025^[update], the family includes these genera:^[1]

• Austrodomus Lawrence, 1947 – southern Africa
• Brasilomma Brescovit, Ferreira & Rheims, 2012 – Brazil
• Caudalia Alayón, 1980 – Cuba
• Chileomma Platnick, Shadab & Sorkin, 2005 – Chile
• Chileuma Platnick, Shadab & Sorkin, 2005 – Chile
• Chilongius Platnick, Shadab & Sorkin, 2005 – Chile
• Eleleis Simon, 1893 – southern Africa
• Indiani Rodrigues, Cizauskas & Lemos, 2020 – Brazil
• Lygromma Simon, 1893 – Costa Rica to Brazil, Galapagos
• Lygrommatoides Strand, 1918 – Japan
• Moreno Mello-Leitão, 1940 – Argentina, Chile
• Namundra Platnick & Bird, 2007 – Angola, Namibia, South Africa
• Neozimiris Simon, 1903 – Bahamas, Colombia, Curaçao, Galapagos, Mexico, Panama, Puerto Rico, United States, Mexico
• Nopyllus Ott, 2014 – Brazil
• Paleotoca Cizauskas, Zampaulo & Brescovit, 2024 – Brazil
• Paracymbiomma Rodrigues, Cizauskas & Rheims, 2018 – Brazil
• Plutonodomus Cooke, 1964 – Tanzania
• Prodidomus Hentz, 1847 – Mediterranean, Africa, Australia, Asia, Venezuela, Hawaii
• Purcelliana Cooke, 1964 – Namibia, South Africa
• Theuma Simon, 1893 – Africa, Turkmenistan (?)
• Tivodrassus Chamberlin & Ivie, 1936 – Mexico
• Tricongius Simon, 1893 – Argentina, Brazil, Colombia, Venezuela
• Zimirina Dalmas, 1919 – North Africa, southern Europe, South Africa
• Zimiris Simon, 1882 – Yemen (Socotra), India. Introduced circumtropical

A 2020 phylogenetic analysis involving 59 species of Prodidominae and 32 outgroup species did not recover Prodidominae as monophyletic because Anagrina did not arise within the subfamily.^[4] The study re-established Molycriinae (including genera Cryptoerithus, Molycria, Nomindra, Wesmaldra, and Wydundra) as a distinct subfamily in Gnaphosidae, a sister to Prodidominae.^[4]