Pupinidae

Pupinidae is a taxonomic family of land snails with an operculum, terrestrial gastropod mollusks in the superfamily Cyclophoroidea (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).^[2]

Pupinidae Temporal range: Cenomanian–Recent PreꞒ Ꞓ O S D C P T J K Pg N
Pupinella rufa
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
Subclass:	Caenogastropoda
Order:	Architaenioglossa
Superfamily:	Cyclophoroidea
Family:	Pupinidae L. Pfeiffer, 1853
Diversity[1]
at least 121 extant species

Distribution

The distribution of the family Pupinidae includes the Himalayas, Assam, Myanmar, peninsular Malaysia, Sumatra, Borneo, Philippines, Vietnam, Thailand,^[3] Australia,^[4] Melanesia, Micronesia and Papua New Guinea.^[5]

Taxonomy

† Coptocheilus electrothauma holotype in Burmese amber

Subfamilies and genera within the family Pupinidae include (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005):^[6]

Pupininae L. Pfeiffer, 1853

• Alpinipupina Stanisic, 2010
• Barnaia Thach, 2017
• Callianella R. B. Newton, 1891^[1]
• Cordillerapina Stanisic, 2010
• Hargravesia H. Adams, 1871
• Hildapina Iredale, 1940
• Ischurostoma Bourguignat, 1874
• Moulinsia Grateloup, 1840^[1]
• Necopupina Iredale, 1937
• Porocallia Möllendorff, 1893
• Pupina Vingard, 1829 - type genus of the subfamily Pupininae^[1][6]
• Signepupina Iredale, 1937
• Siphonostyla Kobelt, 1897
• Tylotoechus Kobelt & Möllendorff, 1897

Liareinae Powell, 1946 - synonym: Cytoidae Climo, 1969 (n.a.) = not available name

Main article: Liareinae

Pollicariinae Thiele, 1929

• Pollicaria Gould, 1856^[1]
• Hybocystis Benson, 1859: synonym of Pollicaria A.A. Gould, 1856

Pupinellinae Kobelt, 1902 - synonyms: Ventriculidae Wenz, 1915, Pollicariini Thiele, 1929

• Barnaia Thach, 2017
• Bellardiella Tapparone Canefri, 1883
• Coptocheilus Gould, 1862
• Csomapupa Páll-Gergely, 2015
• Didomasta Iredale, 1941
• Hedleya Cox, 1892
• † Kallomastoma Stache, 1889
• Nodopomatias Gude, 1921
• Pseudopomatias Möllendorff, 1885^[1]
• Pupinella Gray [in Baird], 1850 - type genus of the subfamily Pupinellinae^[1][6]
• Raphaulus Pfeiffer, 1856^[1]
• Scaeopupina Iredale, 1941
• Schistoloma Kobelt, 1902:^[1][3] synonym of Coptocheilus A. Gould, 1862 (unnecessary replacement name)
• Streptaulus Benson, 1857^[1]
• Tortulosa Gray, 1847^[1]
• Vargapupa Páll-Gergely, 2015

subfamily incertae sedis

• Suavocallia Iredale, 1933^[7]
• (extinct) Cretatortulosa Yu, Wang and Pan, 2018^[8] Burmese amber, Myanmar, Cenomanian.

Ecology

These snails live in wet forests in leaf litter.^[5]