Bayan Mandahu Formation

"Wulansuhai Formation" redirects here. For the Turonian formation, see Ulansuhai Formation.

The Bayan Mandahu Formation (also known as Wulansuhai Formation or Wuliangsuhai Formation) is a geological unit of "redbeds" located near the village of Bayan Mandahu in Inner Mongolia, China, in the Gobi Desert. It dates from the late Cretaceous Period. Laid down in the Campanian, it is dated somewhat uncertainly to between 75 and 71 mya (million years ago).^[1]

Bayan Mandahu Formation
Stratigraphic range: Campanian, 75–71 Ma PreꞒ Ꞓ O S D C P T J K Pg N ↓
Type	Geological formation
Unit of	Gobi Desert
Lithology
Primary	Sandstone
Location
Coordinates	41.75N 106.75E
Region	Inner Mongolia
Country	China
Type section
Named for	Bayan Mandahu, Urad Rear Banner, Inner Mongolia

Description

The paleoenvironment it preserves was semi-arid and characterized by alluvial (stream-deposited) and eolian (wind-deposited) sediments. The formation is known for its vertebrate fossils, most of which are preserved in unstructured sandstone, indicating burial by wind-blown sandstorms.^[2]

Paleofauna of the Bayan Mandahu Formation

The fauna of the Bayan Mandahu is very similar in composition to the nearby Djadochta Formation, and the two may have been deposited at roughly the same time. These two formations share many of the same genera, but differ in the makeup of species. For example, the most common mammal in the Djadochta is Kryptobaatar dashzevegi, while in the Bayan Mandahu, it is the closely related Kryptobaatar mandahuensis. Similarly, the dinosaur fauna of the Djadochta includes Protoceratops andrewsi and Velociraptor mongoliensis, while the Bayan Mandahu contains Protoceratops hellenikorhinus and Velociraptor osmolskae.^[1]

Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon		Notes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.

Crocodylomorphs

Crocodylomorphs of the Bayan Mandahu Formation
Genus	Species	Location	Material	Notes	Images
Shamosuchus[2]				A mesoeucrocodylian.

Lizards

An amphisbaenian species is known from the formation.^[2] An iguanian species is known from the formation.^[2]

Lizards of the Bayan Mandahu Formation
Genus	Species	Location	Material	Notes	Images
Adamisaurus[2]	A. magnidentatus[2]			An agamid.
Bainguis[2]				A diploglossan.
Telmasaurus[2]				A monitor lizard.

Turtles

Turtles of the Bayan Mandahu Formation
Genus	Species	Location	Material	Notes	Images
Basilemys[2]				A softshell turtle.
"Zangerlia"	"Z." neimongolensis	Bayan Mandahu	"Partial to complete skeletons of multiple individuals."[3]	A nanhsiungchelyid.

Mammals

A taeniolabidoidea multituberculate is known from the formation.^[2]

Mammals of the Bayan Mandahu Formation
Genus	Species	Location	Material	Notes	Images
Guibaatar[4]	G. castellanus			A djadochtatheriid multituberculate.
Kennalestes[2]	K. gobiensis[2]			A placental. Also present in the Djadochta Formation.
Kryptobaatar[1]	K. mandahuensis			A multituberculate.

Dinosaurs

Alvarezsaurs

Genus	Species	Location	Material	Notes	Images
Alvarezsauridae indet.[5]	Indeterminate[5]	Bayan Mandahu[5]	An articulated series of partial anterior cervical vertebrae, an anterior cervical vertebra, isolated fragmentary anterior caudal vertebrae, a partial scapula, and pedal phalanges.[5]	An unnamed parvicursorine alvarezsaurid.[5]
Linhenykus	L. monodactylus[6]	Wuliangsuhai [6]	Cervical vertebrae, dorsal vertebrae, sacral vertebrae, caudal vertebrae, a scapulocoracoid, a sternum, much of the forelimbs, a partial pelvis, nearly complete hindlimbs, and some unidentified fragments.[6]	A monodactyl parvicursorine alvarezsaurid.[6]

Ankylosaurs

Genus	Species	Location	Material	Notes	Images
Pinacosaurus	P. grangeri[7]	Bayan Mandahu[7]	[Four] juvenile specimens consisting of skulls, manidibles, cervical half-rings and almost complete skeletons, and an incomplete subadult skull.[7]	A basal ankylosaurine ankylosaurid also known from the Djadochta Formation.
	P. mephistocephalus[8]	Bayan Mandahu[8]	A nearly complete articulated skeleton, with in situ cervical dermal armour and tail-club, that lies in a natural with limb bones doubled under the body.[8]	A basal ankylosaurine ankylosaurid.

Birds

Genus	Species	Location	Material	Notes	Images
cf. Gobipteryx	Indeterminate		"Eggs similar to those attributed to Gobipteryx minuta from the Djadochta Formation."[2]

Ceratopsians

Genus	Species	Location	Material	Notes	Images
Genus	Species	Location	Material	Notes	Images
Bagaceratops	B. rozhdestvenskyi	Bayan Mandahu	A partial skull.[9]	A protoceratopsid also known from the Barun Goyot Formation and possibly the Djadochta Formation.
Magnirostris[1]	M. dodsoni[1]	Bayan Mandahu[10]	A partial skull.	A protoceratopsid now considered synonym of Bagaceratops.[9]
Protoceratops	P. hellenikorhinus[11]	Bayan Mandahu[11]	Skulls and sparse body remains from multiple individuals.[12][11]	A protoceratopsid.
Udanoceratops?	Indeterminate	Bayan Mandahu	"Nearly 1 meter long skull."[2]	A giant leptoceratopsid. Reported remains probably represent Protoceratops hellenikorhinus.[13]

Dromaeosaurs

Genus	Species	Location	Material	Notes	Images
Linheraptor[14]	L. exquisitus[14]	Wuliangsuhai[14]	A skull, cervical vertebrae, caudal vertebrae, a scapula, a humerus, a radius, metacarpals, manual phalanxes, sternal plate, a pubis, a femur, a tibiotarsus, metatarsals, and pedal phalanxes.[14]	A possible velociraptorine dromaeosaurid.[14]
Velociraptor	V. osmolskae[15]	Bayan Mandahu[1]	A paired maxillae and left lacrimal.[1]	A velociraptorine dromaeosaurid.[1]

Hadrosaurs

Genus	Species	Location	Material	Notes	Images
Undescribed Hadrosauridae[10]	Indeterminate[10]	Bayan Mandahu[10]		A hadrosaurid.

Oviraptorosaurs

Genus	Species	Location	Material	Notes	Images
Machairasaurus	M. leptonychus[10]	Bayan Mandahu[10]	A nearly complete articulated right forearm and manus, pedal phalanges, manual phalanges, caudal vertebrae, dorsal ribs, chevrons, and fragments of metatarsals III and IV.[10]	A heyuanniine oviraptorid.[10]
Oviraptoridae indet.	Indeterminate	Bayan Mandahu	"Partial individual sitting atop an egg clutch."[16]	An oviraptorid that represent the second nesting oviraptorid. Sometimes referred to Machairasaurus.[10][17]
Wulatelong	W. gobiensis[18]	Wuliangsuhai[18]	A partial skull, dorsal vertebrae, caudal vertebrae, partial scapulocoracoids, a partial sternum, a partial humerus, an incomplete manus, a nearly complete pelvic girdle, and the right hindlimb.[18]	A basal oviraptorid oviraptorosaur.[18]

Sauropods

Genus	Species	Location	Material	Notes	Images
Undescribed Sauropoda[10]	Indeterminate[10]	Bayan Mandahu[10]		A sauropod.

Troodontids

Genus	Species	Location	Material	Notes	Images
Linhevenator[19]	L. tani[19]	Wuliangsuhai[19]	A skull and mandible, anterior dorsal vertebrae, middle dorsal vertebrae, a scapula, a humerus, incomplete ischia, a femur, nearly complete pes, and other fragmentary bones.[19]	A troodontid with short forearms.[19]
Papiliovenator	P. neimengguensis[20]	Bayan Mandahu[20]	An almost complete skull, cervical vertebrae, anterior dorsal vertebrae, partial scapulae, partial coracoid, fragmentary pelvic girdle, partial humeri, ulna, radius, partial manus, femur, tibia, fibula, pes, and a pedal phalanx II-3.[20]	A troodontid that is part of a clade that includes all other Late Cretaceous troodontids except Almas.[20]
Philovenator	P. curriei[21]	Wuliangsuhai[21]	A nearly complete femur, a partial tibia, a partial fibula, astragalus-calcaneum complex, distal tarsals, metatarsals, and pedal phalanges.[21]	A troodontid.[21]

Tyrannosaurs

Genus	Species	Location	Material	Notes	Images
Undescribed Tyrannosauridae[10]	Indeterminate[10]	Bayan Mandahu[10]		A tyrannosaurid.

See also

• China portal
• Paleontology portal

• Barun Goyot Formation
• List of fossil sites (with link directory)
• Nemegt Formation
• List of dinosaur-bearing rock formations