Carol Greider

Carolyn Widney "Carol" Greider (born April 15, 1961) is an American molecular biologist. She is Professor and Director of Molecular Biology and Genetics at Johns Hopkins University.^[1]

Carol Greider
Born	(1961-04-15) April 15, 1961 (age 64) San Diego, California
Nationality	American
Alma mater	University of California, Santa Barbara University of California, Berkeley
Known for	discovery of telomerase
Awards	Lasker Award (2006) Louisa Gross Horwitz Prize (2007) Nobel Prize in Physiology or Medicine (2009)
Scientific career
Fields	Molecular biology
Institutions	Cold Spring Harbor Laboratory Johns Hopkins University
Doctoral advisor	Elizabeth Blackburn

Greider discovered the enzyme telomerase in 1984, when she was a graduate student of Elizabeth Blackburn at the University of California, Berkeley. Greider pioneered research on the structure of telomeres, the ends of the chromosomes. She was awarded the 2009 Nobel Prize for Physiology or Medicine, with Blackburn and Jack Szostak, for their discovery that telomeres are protected from progressive shortening by the enzyme telomerase.^[2]