竹叶青蛇属

竹叶青蛇属
	白唇竹叶青（Trimeresurus albolabris）
科学分类
界：	动物界 Animalia
门：	脊索动物门 Chordata
纲：	爬行纲 Reptilia
目：	有鳞目 Squamata
亚目：	蛇亚目 Serpentes
科：	蝰科 Viperidae
亚科：	蝮蛇亚科 Crotalinae
属：	竹叶青蛇属 Trimeresurus; Lacépède, 1804
种
	约30种，详见内文。
异名
	Cryptelytrops Malhotra & Thorpe, 2004; Parias Gray, 1849; Peltopelor Vogel et al., 2004; Viridovipera Malhotra & Thorpe, 2004; Craspedocephalus ; Kuhl & van Hasselt, 1822; Trimeresura – Fleming, 1822; Craspedocephalus – Gray, 1825; Megaera Wagler, 1830; Atropos Wagler, 1830; Trimesurus Gray, 1842;

竹叶青属（学名：Trimeresurus），又名竹叶青蛇属、青竹丝属，是蝰蛇科的其中一属，主要分布在亚洲的印度、东南亚、中国以及太平洋上的岛屿，共有三十多种，台湾话称为青竹丝，属于破坏血小板凝固功能的出血性毒蛇。

Quick Facts 竹叶青蛇属, 科学分类 ...

Close

特征

竹叶青蛇是具管牙、颊窝的毒蛇。牙较大，但没有蝰蛇牙大。头三角形，吻尖。尾巴后面红了一节，这是与翠青蛇最明显的区别。南亚采茶时传职业伤害被此蛇咬到手指。

血循毒为主，平均一次排毒5毫克。人致死剂量100毫克。

分类及分布

广泛分布于除北亚、中亚、西亚和日本以外所有的亚洲，常见攀附在竹丛上。

截止2022年6月1日，根据reptile-database中的数据，本属包含43种：^[1]

白唇竹叶青 Trimeresurus albolabris (Gray, 1842)
Trimeresurus andalasensis David, Vogel, Vijayakumar & Vidal 2006^[2]
安氏竹叶青 Trimeresurus andersonii (Theobald, 1868)
藏南竹叶青蛇 Trimeresurus arunachalensi Captain, Deepak, Pandit, Bhatt & Athreya, 2019
Trimeresurus ayeyarwadyensis Chan, Anuar, Sankar, I.T. Law, I.S. Law, Shivaram, Christian, Mulcahy & Malhotra, 2023^[3]
Trimeresurus buniana Grismer, Grismer & McGuire 2006^[4]
Trimeresurus calamitas Vogel, David & Sidik, 2022^[5]
堪氏竹叶青 Trimeresurus cantori (Blyth, 1846)
豆蔻竹叶青 Trimeresurus cardamomensis (Malhotra, Thorpe, Mrinalini & Stuart, 2011)^[6]
饰尾竹叶青 Trimeresurus caudornatus Chen，Ding，Vogel & Shi, 2020
Trimeresurus ciliaris Idiiatullina, Pawangkhanant, Tawan, Worranuch, Dechochai, Suwannapoom, Nguyen, Chanhome & Poyarkov, 2023^[7]
Trimeresurus cyanolabris Idiiatullina, Nguyen, Bragin, Pawangkhanant, Le, Vogel, David & Poyarkov, 2024 ^[8]
戴氏竹叶青 Trimeresurus davidi Chandramouli, Campbell & Vogel, 2020
缅甸竹叶青 Trimeresurus erythrurus (Cantor, 1839)
条纹竹叶青 Trimeresurus fasciatus （Boulenger, 1896)
菲律宾竹叶青 Trimeresurus flavomaculatus (Gray, 1842)
台湾竹叶青蛇 Trimeresurus gracilis Oshima, 1920，台湾竹叶青
冈氏竹叶青 Trimeresurus gumprechti (David, Vogel, Pauwels & Vidal, 2002)
古氏竹叶青 Trimeresurus gunaleni Vogel, David & Sidik, 2014
滇南竹叶青 Trimeresurus guoi Chen, Shi, Vogel & Ding, 2021^[9]
哈氏竹叶青 Trimeresurus hageni (Lidth De Jeude, 1886)
鸿山竹叶青 Trimeresurus honsonensis (Grismer, Ngo & Grismer, 2008)
巽他竹叶青 Trimeresurus insularis Kramer, 1977
坎布里竹叶青 Trimeresurus kanburiensis (Smith, 1943)
Trimeresurus kirscheyi Vogel, David & Sidik, 2022^[5]
Trimeresurus kraensis Idiiatullina, Pawangkhanant, Suwannapoom, Tawan, Chanhome, Nguyen, David, Vogel & Poyarkov, 2024^[10]
圭布里竹叶青 Trimeresurus kuiburi Sumontha, Suntrarachun, Pauwels, Pawangkhanant, Chomngam, Iamwiriyakul & Chanhome, 2021
尼科巴竹叶青 Trimeresurus labialis (Steindachner, 1867)
Trimeresurus lanna Idiiatullina, Nguyen, Pawangkhanant, Suwannapoom, Chanhome, Mirza, David, Vogel & Poyarkov, 2024^[11]
大眼竹叶青 Trimeresurus macrops (Kramer, 1977)^[6]
马氏竹叶青 Trimeresurus malcolmi (Loveridge, 1938)
玛亚竹叶青 Trimeresurus mayaae Rathee, Purkayastha, Lalremsanga, Dalal, Biakzuala, Muansanga & Mirza, 2022^[12]
麦氏竹叶青 Trimeresurus mcgregori Taylor, 1919
墨脱竹叶青蛇 Trimeresurus medoensis (Zhao, 1977)，墨脱竹叶青
格莫尔达竹叶青 Trimeresurus mutabilis Stoliczka, 1870
云斑竹叶青 Trimeresurus nebularis Vogel, David & Pauwels, 2004
普吉竹叶青 Trimeresurus phuketensis Sumontha, Kunya, Pauwels, Nitikul & Punnadee, 2011
坡普竹叶青蛇 Trimeresurus popeiorum Smith, 1937
Trimeresurus puniceus Boie, 1827^[2]，为 Craspedocephalus puniceus (Boie, 1827)的异名
红树竹叶青 Trimeresurus purpureomaculatus (Gray in Gray & Hardwicke, 1832)
红眼竹叶青 Trimeresurus rubeus (Malhotra, Thorpe, Mrinalini & Stuart, 2011)
沙巴竹叶青 Trimeresurus sabahi Regenass & Kramer, 1981
萨氏竹叶青蛇 Trimeresurus salazar Mirza, Bhosale, Phansalkar, Sawant, Gowande & Patel, 2020
舒氏竹叶青 Trimeresurus schultzei (Griffin, 1909)
克拉竹叶青 Trimeresurus septentrionalis (Kramer, 1977)
四川竹叶青 Trimeresurus sichuanensis (Guo & Wang, 2011)
赤尾青竹丝 Trimeresurus stejnegeri (Schmidt, 1925)^[13]
苏门答腊竹叶青 Trimeresurus sumatranus (Raffles, 1822)^[14]
西藏竹叶青 Trimeresurus tibetanus Huang, 1982
越南竹叶青 Trimeresurus truongsonensis Orlov, Ryabov, Thanh & H Cuc, 2004
Trimeresurus uetzi ^[15]
泰国竹叶青 Trimeresurus venustus (Vogel, 1991)
沃氏竹叶青 Trimeresurus vogeli (David, Vidal & Pauwels, 2001)
Trimeresurus whitteni Vogel, David & Sidik, 2022^[5]
Trimeresurus wiroti Trutnau, 1981 ^[2]
盈江竹叶青蛇 Trimeresurus yingjiangensis Chen, Zhang, Shi, Tang, Guo, Song & Ding, 2019
云南竹叶青 Trimeresurus yunnanensis (Schmidt, 1925)

2004年，英国班戈大学的阿妮塔·马尔霍特拉（Anita Malhotra）和罗杰·S·索普（Roger S Thorpe）结合半阴茎形态、鳞片特征和分子生物学证据，对广义的竹叶青属进行了系统学研究，认为广义的竹叶青属不是一个单系，恢复腺蝮属（Cryptelytrops）、等蝮属（Parias）和奇盾蝮属（Peltopelor）3属，新建了喜山蝮属（Himalayophis）、坡普蝮属（Popeia）和绿蝮属（Viridovipera）3属。^[16]^[17]但是也有学者认为被拆分的诸属实际上是竹叶青属的亚属。^[18]

参考文献

[1]
Reptarium.cz Reptile Database 中的 Trimeresurus
[2]
David, Patrick; Vogel, Gernot; Vijayakumar, S.P.; Vidal, Nicolas. 2006. A revision of the Trimeresurus puniceus – complex (Serpentes: Viperidae: Crotalinae) based on morphological and molecular data. Zootaxa. 1293:1-78
[3]
Kin Onn Chan, Shahrul Anuar, Ananthanarayanan Sankar, Ingg Thong Law, Ing Sind Law, Rasu Shivaram, Ching Christian, Daniel G. Mulcahy and Anita Malhotra. 2023. A New Species of Pit-viper from the Ayeyarwady and Yangon regions in Myanmar (Viperidae, Trimeresurus). ZooKeys. 1186: 221-234. DOI: 10.3897/zookeys.1186.110422
[4]
Grismer L. L., Grismer J. L. & McGuire J. A., 2006. A new species of pitviper of the genus Popeia (Squamata: Viperidae) from Pulau Tioman, Pahang, West Malaysia. Zootaxa. 1305: 1-19.
[5]
Gernot Vogel, Patrick David and Irvan Sidik. 2022. A Review of the Complex of Trimeresurus hageni (Lidth de Jeude, 1886) (Squamata: Viperidae) with Descriptions of Three New Insular Species from Indonesia. TAPROBANICA. 11(2); 54–83. DOI: 10.47605/tapro.v11i2.283
[6]
Malhotra, A., R. S. Thorpe, Mrinalini and B. L. Stuart. 2011. Two new species of pitviper of the genus Cryptelytrops Cope 1860 (Squamata: Viperidae: Crotalinae) from Southeast Asia. Zootaxa. 2757: 1–23. http://www.mapress.com/zootaxa/2011/f/zt02757p023.pdf （页面存档备份，存于互联网档案馆）
[7]
Sabira S. Idiiatullina, Parinya Pawangkhanant, Tanapong Tawan, Thanawut Worranuch, Bunyarit Dechochai, Chatmongkon Suwannapoom, Tan Van Nguyen, Lawan Chanhome and Nikolay A. Poyarkov. 2023. Limestone Jewel: A New colourful Karst-dwelling Pitviper (Serpentes: Viperidae: Trimeresurus) from the poorly explored borderlands of southern peninsular Thailand. Vertebrate Zoology. 73: 697-716. DOI: 10.3897/vz.73.e109854
[8]
Sabira S. Idiiatullina, Tan Van Nguyen, Andrey M. Bragin, Parinya Pawangkhanant, Dac Xuan Le, Gernot Vogel, Patrick David and Nikolay A. Poyarkov. 2024. A New Species of Green Pitviper of the Trimeresurus macrops complex (Reptilia: Serpentes: Viperidae) from South Central Coastal Region of Vietnam. Zootaxa. 5474(4); 375-411. DOI: doi.org/10.11646/zootaxa.5474.4.3
[9]
Zening CHEN, Shengchao SHI, Jun GAO, Gernot VOGEL, Zhaobin SONG, Li DING and Rong DAI. 2021. A New Species of Trimeresurus Lacépède, 1804 (Squamata: Viperidae) from Southwestern China, Vietnam, Thailand and Myanmar. Asian Herpetological Research. 11(4); 1-11. DOI: 10.16373/j.cnki.ahr.200084
[10]
Sabira S. Idiiatullina; Parinya Pawangkhanant, Chatmongkon Suwannapoom, Tanapong Tawan, Lawan Chanhome, Tan Van Nguyen, Patrick David, Gernot Vogel and Nikolay A. Poyarkov. 2024. Another New Species of Karst-associated Pitviper (Serpentes, Viperidae: Trimeresurus) from the Isthmus of Kra, Peninsular Thailand. European Journal of Taxonomy. 930(1), 20–52. DOI: 10.5852/ejt.2024.930.2489
[11]
Sabira S. Idiiatullina, Tan Van Nguyen, Parinya Pawangkhanant, Chatmongkon Suwannapoom, Lawan Chanhome, Zeeshan A. Mirza, Patrick David, Gernot Vogel and Nikolay A. Poyarkov. 2024. An Integrative Taxonomic Revision of the Trimeresurus popeiorum group of Pitvipers (Reptilia: Serpentes: Viperidae) with Descriptions of Two New Species from the Indo-Burma Biodiversity Hotspot. Vertebrate Zoology. 74: 303-342. DOI: 10.3897/vz.74.e113347
[12]
Yashpal Singh Rathee, Jayaditya Purkayastha, Hmar Tlawmte Lalremsanga, Siddharth Dalal, Lal Biakzuala, Lal Muansanga and Zeeshan A. Mirza. 2022. A New Cryptic Species of Green Pit Viper of the Genus Trimeresurus Lacépède, 1804 (Serpentes, Viperidae) from northeast India. PLoS ONE. 17(5): e0268402. DOI: 10.1371/journal.pone.0268402
[13]
Laura Rodrigues Vieira de Alencar, Marcio Martins, Gustavo Burin and Tiago Bosisio Quental. 2017. Arboreality Constrains Morphological Evolution but Not Species Diversification in Vipers. Proceedings of the Royal Society B: Biological Sciences. 284(1869) DOI: 10.1098/rspb.2017.1775
[14]
Gernot Vogel, Patrick David and Irvan Sidik. 2014. On Trimeresurus sumatranus (Raffles, 1822), with the Designation of A Neotype and the Description of A New Species of Pitviper from Sumatra (Squamata: Viperidae: Crotalinae). Amphibian & Reptile Conservation 8(2) [General Issue]: 1–29 (e80).
[15]
Gernot Vogel, Tan Van Nguyen, Patrick David. 2023. A New Green Pitviper of the Trimeresurus albolabris complex (Reptilia, Serpentes, Viperidae) from central and southern Myanmar. Zootaxa. 5357(4); 515-554. DOI: 10.11646/zootaxa.5357.4.3
[16]
郭鹏. 广义竹叶青蛇属的分类及系统发育研究(蛇亚目:蝮亚科)[D]. 四川大学. 2005 [2021-10-24]. doi:10.7666/d.y775610. （原始内容存档于2021-10-26）.
[17]
Malhotra A, Thorpe RS. A phylogeny of four mitochondrial gene regions suggests a revised taxonomy for Asian pitvipers (Trimeresurus and Ovophis). Molecular Phylogenetics and Evolution. 2004, 32: 83–100.
[18]
P David，G Vogel，A Dubois. On the need to follow rigorously the Rules of the Code for the subsequent designation of a nucleospecies (type species) for a nominal genus which lacked one: the case of the nominal genus Trimeresurus Lacepede, 1804 (Reptilia: Squamata: Viperidae). Zootaxa. 2011, 2992: 1–51 [2021-10-24]. doi:10.11646/zootaxa.2992.1.1. （原始内容存档于2021-10-24）.

[RDB-1] [1]
Reptarium.cz Reptile Database 中的 Trimeresurus

[未命名-20240110220100-2] [2]
David, Patrick; Vogel, Gernot; Vijayakumar, S.P.; Vidal, Nicolas. 2006. A revision of the Trimeresurus puniceus – complex (Serpentes: Viperidae: Crotalinae) based on morphological and molecular data. Zootaxa. 1293:1-78

[3] [3]
Kin Onn Chan, Shahrul Anuar, Ananthanarayanan Sankar, Ingg Thong Law, Ing Sind Law, Rasu Shivaram, Ching Christian, Daniel G. Mulcahy and Anita Malhotra. 2023. A New Species of Pit-viper from the Ayeyarwady and Yangon regions in Myanmar (Viperidae, Trimeresurus). ZooKeys. 1186: 221-234. DOI: 10.3897/zookeys.1186.110422

[4] [4]
Grismer L. L., Grismer J. L. & McGuire J. A., 2006. A new species of pitviper of the genus Popeia (Squamata: Viperidae) from Pulau Tioman, Pahang, West Malaysia. Zootaxa. 1305: 1-19.

[Vogel-5] [5]
Gernot Vogel, Patrick David and Irvan Sidik. 2022. A Review of the Complex of Trimeresurus hageni (Lidth de Jeude, 1886) (Squamata: Viperidae) with Descriptions of Three New Insular Species from Indonesia. TAPROBANICA. 11(2); 54–83. DOI: 10.47605/tapro.v11i2.283

[未命名_2-20240110220100-6] [6]
Malhotra, A., R. S. Thorpe, Mrinalini and B. L. Stuart. 2011. Two new species of pitviper of the genus Cryptelytrops Cope 1860 (Squamata: Viperidae: Crotalinae) from Southeast Asia. Zootaxa. 2757: 1–23. http://www.mapress.com/zootaxa/2011/f/zt02757p023.pdf （页面存档备份，存于互联网档案馆）

[7] [7]
Sabira S. Idiiatullina, Parinya Pawangkhanant, Tanapong Tawan, Thanawut Worranuch, Bunyarit Dechochai, Chatmongkon Suwannapoom, Tan Van Nguyen, Lawan Chanhome and Nikolay A. Poyarkov. 2023. Limestone Jewel: A New colourful Karst-dwelling Pitviper (Serpentes: Viperidae: Trimeresurus) from the poorly explored borderlands of southern peninsular Thailand. Vertebrate Zoology. 73: 697-716. DOI: 10.3897/vz.73.e109854

[8] [8]
Sabira S. Idiiatullina, Tan Van Nguyen, Andrey M. Bragin, Parinya Pawangkhanant, Dac Xuan Le, Gernot Vogel, Patrick David and Nikolay A. Poyarkov. 2024. A New Species of Green Pitviper of the Trimeresurus macrops complex (Reptilia: Serpentes: Viperidae) from South Central Coastal Region of Vietnam. Zootaxa. 5474(4); 375-411. DOI: doi.org/10.11646/zootaxa.5474.4.3

[9] [9]
Zening CHEN, Shengchao SHI, Jun GAO, Gernot VOGEL, Zhaobin SONG, Li DING and Rong DAI. 2021. A New Species of Trimeresurus Lacépède, 1804 (Squamata: Viperidae) from Southwestern China, Vietnam, Thailand and Myanmar. Asian Herpetological Research. 11(4); 1-11. DOI: 10.16373/j.cnki.ahr.200084

[10] [10]
Sabira S. Idiiatullina; Parinya Pawangkhanant, Chatmongkon Suwannapoom, Tanapong Tawan, Lawan Chanhome, Tan Van Nguyen, Patrick David, Gernot Vogel and Nikolay A. Poyarkov. 2024. Another New Species of Karst-associated Pitviper (Serpentes, Viperidae: Trimeresurus) from the Isthmus of Kra, Peninsular Thailand. European Journal of Taxonomy. 930(1), 20–52. DOI: 10.5852/ejt.2024.930.2489

[11] [11]
Sabira S. Idiiatullina, Tan Van Nguyen, Parinya Pawangkhanant, Chatmongkon Suwannapoom, Lawan Chanhome, Zeeshan A. Mirza, Patrick David, Gernot Vogel and Nikolay A. Poyarkov. 2024. An Integrative Taxonomic Revision of the Trimeresurus popeiorum group of Pitvipers (Reptilia: Serpentes: Viperidae) with Descriptions of Two New Species from the Indo-Burma Biodiversity Hotspot. Vertebrate Zoology. 74: 303-342. DOI: 10.3897/vz.74.e113347

[12] [12]
Yashpal Singh Rathee, Jayaditya Purkayastha, Hmar Tlawmte Lalremsanga, Siddharth Dalal, Lal Biakzuala, Lal Muansanga and Zeeshan A. Mirza. 2022. A New Cryptic Species of Green Pit Viper of the Genus Trimeresurus Lacépède, 1804 (Serpentes, Viperidae) from northeast India. PLoS ONE. 17(5): e0268402. DOI: 10.1371/journal.pone.0268402

[13] [13]
Laura Rodrigues Vieira de Alencar, Marcio Martins, Gustavo Burin and Tiago Bosisio Quental. 2017. Arboreality Constrains Morphological Evolution but Not Species Diversification in Vipers. Proceedings of the Royal Society B: Biological Sciences. 284(1869) DOI: 10.1098/rspb.2017.1775

[14] [14]
Gernot Vogel, Patrick David and Irvan Sidik. 2014. On Trimeresurus sumatranus (Raffles, 1822), with the Designation of A Neotype and the Description of A New Species of Pitviper from Sumatra (Squamata: Viperidae: Crotalinae). Amphibian & Reptile Conservation 8(2) [General Issue]: 1–29 (e80).

[15] [15]
Gernot Vogel, Tan Van Nguyen, Patrick David. 2023. A New Green Pitviper of the Trimeresurus albolabris complex (Reptilia, Serpentes, Viperidae) from central and southern Myanmar. Zootaxa. 5357(4); 515-554. DOI: 10.11646/zootaxa.5357.4.3

[16] [16]
郭鹏. 广义竹叶青蛇属的分类及系统发育研究(蛇亚目:蝮亚科)[D]. 四川大学. 2005 [2021-10-24]. doi:10.7666/d.y775610. （原始内容存档于2021-10-26）.

[17] [17]
Malhotra A, Thorpe RS. A phylogeny of four mitochondrial gene regions suggests a revised taxonomy for Asian pitvipers (Trimeresurus and Ovophis). Molecular Phylogenetics and Evolution. 2004, 32: 83–100.

[18] [18]
P David，G Vogel，A Dubois. On the need to follow rigorously the Rules of the Code for the subsequent designation of a nucleospecies (type species) for a nominal genus which lacked one: the case of the nominal genus Trimeresurus Lacepede, 1804 (Reptilia: Squamata: Viperidae). Zootaxa. 2011, 2992: 1–51 [2021-10-24]. doi:10.11646/zootaxa.2992.1.1. （原始内容存档于2021-10-24）.

[1]

[2]

[3]

[4]

[5]

[6]

[7]

[8]

[9]

[10]

[11]

[12]

[13]

[14]

[15]

[16]

[17]

[18]