2013 in paleobotany

This list of 2013 in paleobotany records new fossil plant taxa that were described during 2013, as well as other significant discoveries and events related to paleobotany that occurred in the year.

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Chlorophyta

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Cymatiosphaera scitula[1]	Sp. nov.	Valid	Wicander & Playford	Devonian (Famennian)	Saverton Shale	United States	A member of Chlorophyta belonging to the group Prasinophyceae.

Bryophyta

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Brachytheciites sachalinensis[2]	Sp. nov	Valid	Ignatov & Perkovsky	Middle Eocene	Sakhalinian amber	Russia	A moss belonging to the family Brachytheciaceae.
Bryiidites[3]	Gen. et sp. nov.	Valid	Barclay et al.	Late Cretaceous (late Cenomanian)	Middle Dakota Formation	United States	A member of Bryidae of uncertain phylogenetic placement. The type specimen is Bryiidites utahensis.
Bryokhutuliinia crassimarginata[4]	Sp. nov	Valid	Ignatov et al.	Late Jurassic		Russia	A moss.
Calymperites[5]	Gen. et sp. nov	Valid	Ignatov & Perkovsky	Late Eocene	Rovno amber	Ukraine	A moss belonging to the group Dicranales (sensu lato). Genus includes new species C. ucrainicus.
Pseudoleskeellites[2]	Gen. et sp. nov	Valid	Ignatov & Perkovsky	Middle Eocene	Sakhalinian amber	Russia	A moss belonging to the order Hypnales. Genus includes new species P. obscurus.

Ferns and fern allies

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Ashicaulis beipiaoensis[6]	Sp. nov	Valid	Tian et al.	Middle Jurassic	Tiaojishan Formation	China	A member of Osmundaceae. Originally described as a species of Ashicaulis; transferred to the genus Millerocaulis by Bomfleur, Grimm & McLoughlin (2017).[7]
Mygdykitia[8]	Gen. et sp. nov	Valid	Golovneva & Shczepetov	Late Cretaceous (Campanian)	Ola Formation	Russia	A fern belonging to the family Dipteridaceae. Genus includes new species M. ochotica.
Palaeosorum[9]	Gen. et sp. nov	Valid	Jacques & Zhou in Jacques, Su & Zhou	Miocene	Dajie Formation	China	A member of Polypodiaceae. The type specimen is Palaeosorum ellipticum.
Todea amissa[10]	Sp. nov.	Valid	Carvalho in Carvalho et al.	Early Eocene		Argentina	A species of Todea.
Yavanna[11]	Gen. et sp. nov	Valid	Vera	Early Cretaceous (Aptian)	Cerro Negro Formation	Antarctica (Livingston Island)	A cyathealean of uncertain placement. Type species is Y. chimaerica.

Ginkgophytes

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Allicospermum angrenicum[12]	Sp. nov.	Valid	Nosova	Middle Jurassic	Angren Formation	Uzbekistan	A Ginkgo-like plant described on the basis of seeds.
Nagrenia[12]	Gen. et sp. nov.	Valid	Nosova	Middle Jurassic	Angren Formation	Uzbekistan	A member of Ginkgoales described on the basis of peduncles furcating into several pedicels. Genus includes new species N. samylinae.

Gnetophyta

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Chengia[13]	Gen. et sp. nov.	Valid	Yang, Lin & Wang	Early Cretaceous (Aptian)	Yixian Formation	China	A member of Ephedraceae. The type specimen is Chengia laxispicata.
Friedsellowia gracilifolia[14]	Gen. nov. et sp. nov.	Valid	Löwe et al.	Early Cretaceous	Crato Formation	Brazil	A gnetophyte. The type of a new genus.
Itajuba[citation needed]	Gen. et sp. nov	Valid	Ricardi-Branco et al.	Early Cretaceous (Aptian or Albian)	Crato Member of the Santana Formation	Brazil	A member of Gnetales. The type specimen is Itajuba yansanae.

Pachytestopsida

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Stephanospermum braidwoodensis[15]	Sp. nov.	Valid	Spencer, Hilton & Sutton	Carboniferous	Mazon Creek fossil beds	United States	A member of Medullosales, a species of Stephanospermum.
Stephanospermum shuichengensis[16]	Sp. nov.	Valid	Spencer et al.	Permian (Lopingian)	Wangjiazhai Formation	China	A member of Medullosales, a species of Stephanospermum.

Pinophyta

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Bancroftiastrobus[17]	Gen. et sp. nov.	Valid	Rothwell et al.	Late Jurassic (early Kimmeridgian)	Kimmeridge Clay Formation	United Kingdom	A pinophyte of uncertain phylogenetic placement. Type species is B. digitata.
Brachyoxylon currumilii[18]	Sp. nov.	Valid	Bodnar et al.	Early–Middle Jurassic	Cañadón Asfalto Formation	Argentina	A member of Pinophyta described on the basis of fossil wood.
Cunninghamia taylorii[19]	Sp. nov.	Valid	Serbet, Bomfleur & Rothwell	Late Cretaceous (Campanian)	Horseshoe Canyon Formation	Canada	A species of Cunninghamia.
Elatocladus anorovii[8]	Sp. nov	Valid	Golovneva & Shczepetov	Late Cretaceous (Campanian)	Ola Formation	Russia	A conifer of uncertain phylogenetic placement.
Pagiophyllum heshanense[20]	Sp. nov.	Valid	Liu & Yao	Permian		China	A species of Pagiophyllum.
Pararaucaria delfueyoi[21]	Sp. nov.	Valid	Escapa et al.	Late Jurassic	Cañadón Calcáreo Formation	Argentina	A member of Pinophyta belonging to the family Cheirolepidiaceae, a species of Pararaucaria.
Picea yanensis[8]	Sp. nov	Valid	Golovneva & Shczepetov	Late Cretaceous (Campanian)	Ola Formation	Russia	A spruce.
Prumnopitys anglica[22]	Sp. nov.	Valid	Greenwood, Hill & Conran	Eocene	Branksome Formation	United Kingdom	A species of Prumnopitys.
Schizolepidopsis canicularis[23]	Sp. nov.	Valid	Leslie et al.	Early Cretaceous (probably Aptian-Albian)	Tevshiin Govi Formation	Mongolia	A member of Pinophyta, possibly a member of the stem group of Pinaceae; a species of Schizolepidopsis.
Xuanweioxylon[24]	Gen. et sp. nov.	Valid	He et al.	Permian (late Wuchiapingian-Changhsingian)	Xuanwei Formation	China	A member of Pinophyta. Genus includes new species X. scalariforme.

Angiosperms

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Anonaspermum warmanensis[25]	Sp. nov.	Valid	Wang, Blanchard & Dilcher	Eocene	Cockfield Formation	United States	A member of Annonaceae.
Atli[26]	Gen. et sp. nov.	Valid	Smith et al.	Late Cretaceous (Campanian)		Canada	A ranunculalean known from a stem with lianoid anatomy. The type specimen is Atli morinii.
Bambusiculmus[27]	Gen. et 2 sp. nov.	Valid	Wang & Zhou in Wang et al.	Miocene	Dajie Formation	China	A bamboo. The type specimen is Bambusiculmus angustus; genus also includes Bambusiculmus latus.
Bambusium angustifolia[27]	Sp. nov.	Valid	Wang & Zhou in Wang et al.	Miocene	Dajie Formation	China	A bamboo, a species of Bambusium.
Bambusium latifolia[27]	Sp. nov.	Valid	Wang & Zhou in Wang et al.	Miocene	Dajie Formation	China	A bamboo, a species of Bambusium.
Bombacacioxylon[28]	Gen. et sp. nov.	Valid	Mehrotra et al.	Late Miocene to early Pliocene	Tipam Group	India	A possible relative of Bombax. The type specimen is Bombacacioxylon tertiarum.
Carpolithus warmanensis[25]	Sp. nov.	Valid	Wang, Blanchard & Dilcher	Eocene	Cockfield Formation	United States	A fossil fruit.
Cornerocarpon[25]	Gen. et sp. nov.	Valid	Grote in Wang, Blanchard & Dilcher	Eocene	Cockfield Formation	United States	A member of Rosales belonging or related to the family Moraceae. Genus includes new species C. copiosum.
Corylopsis grotei[25]	Sp. nov.	Valid	Wang, Blanchard & Dilcher	Eocene	Cockfield Formation	United States	A species of Corylopsis.
Diospyros neyveliensis[29]	Sp. nov.	Valid	Mukherjee & Prasad	Miocene	Neyveli Formation	India	A member of the family Ebenaceae described on the basis of fossil wood. Originally described as a species of Diospyros; Srivastava et al. (2018) transferred this species to the genus Ebenoxylon.[30]
Firmiana oligocenica[31]	Sp. nov.	Valid	Srivastava & Mehrotra	Oligocene (Chattian)	Tikak Parbat Formation	India	A species of Firmiana.
Grotea[25]	Gen. et sp. nov.	Valid	Wang, Blanchard & Dilcher	Eocene	Cockfield Formation	United States	A member of the family Bignoniaceae. Genus includes new species G. warmanensis.
Guadua morronei[32]	Sp. nov.	Valid	Brea, Zucol & Franco	Pliocene–Pleistocene	Ituzaingó Formation	Argentina	A bamboo, a species of Guadua.
Hamamelis warmanensis[25]	Sp. nov.	Valid	Wang, Blanchard & Dilcher	Eocene	Cockfield Formation	United States	A witch-hazel.
Indovitis[33]	Gen. et sp. nov.	Valid	Manchester, Kapgate & Wen	Late Cretaceous (Maastrichtian)	Deccan Intertrappean Beds	India	A member of the family Vitaceae. Genus includes new species I. chitaleyae.
Iterophyllum[34]	Gen. et sp. nov.	Valid	Barral et al.	Early Cretaceous (late Barremian)		Spain	A basal eudicot. The type specimen is Iterophyllum lobatum.
Jaguariba wiersemana[35]	Gen. nov. et sp. nov.	Valid	Coiffard, Mohr & Bernardes-de-Oliveira	Early Cretaceous	Crato Formation	Brazil	Fossil Nymphaeales.
Lagerstroemia corvinusii[36]	Sp. nov.	Valid	Prasad	Mio-Pliocene	Churia Formation	Nepal	A species of Lagerstroemia.
Laurelia otagoensis[37]	Sp. nov	Valid	Conran, Bannister & Lee	Early Miocene		New Zealand	A species of Laurelia.
Lauroflorum[25]	Gen. et sp. nov.	Valid	Wang, Blanchard & Dilcher	Eocene	Cockfield Formation	United States	A member of Lauraceae. Genus includes new species L. warmanensis.
Lindera acuminatissima[38]	Sp. nov	Valid	Dao & Sun in Dao et al.	Late Pliocene	Mangbang Formation	China	A species of Lindera.
"Magnolia" szakmanycsabae[39]	Sp. nov.	Valid	Hably	Late Miocene		Hungary	A plant of uncertain phylogenetic placement, possibly a relative of Magnoliaceae–Lauraceae.
Micropetasos[40]	Gen. et sp. nov	Valid	Poinar, Chambers & Wunderlich	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	An early flowering plant of uncertain phylogenetic placement. Genus includes new species M. burmensis.
Myrtineoxylon[41]	Gen. et sp. nov.	Valid	Oskolski, Feng & Jin	Late Eocene	Youganwo Formation	China	A member of Myrtaceae described on the basis of fossil wood. Genus includes new species M. maomingensis.
Palmoxylon vastanensis[42]	Sp. nov.	Valid	Prasad et al.	Eocene	Cambay Shale Formation	India	A palm tree, a species of Palmoxylon.
Paranephelium makumensis[31]	Sp. nov.	Valid	Srivastava & Mehrotra	Oligocene (Chattian)	Tikak Parbat Formation	India	A species of Paranephelium.
Poeciloneuron preindicum[43]	Sp. nov.	Valid	Srivastava & Mehrotra	Late Oligocene		India	A species of Poeciloneuron.
Pongamia arjunkholaensis[36]	Sp. nov.	Valid	Prasad	Mio-Pliocene	Churia Formation	Nepal	A species of Pongamia.
Potomacapnos[44]	Gen. et sp. nov.	Valid	Jud & Hickey	Early Cretaceous (Aptian)	Potomac Group	United States	A close relative of ranunculalean eudicots. The type specimen is Potomacapnos apeleutheron.
Pterygota palaeoalata[31]	Sp. nov.	Valid	Srivastava & Mehrotra	Oligocene (Chattian)	Tikak Parbat Formation	India	A species of Pterygota.
Sapindus palaeoemarginatus[31]	Sp. nov.	Valid	Srivastava & Mehrotra	Oligocene (Chattian)	Tikak Parbat Formation	India	A species of Sapindus.
Spixiarum kipea[45]	Gen. nov. et sp. nov.	Valid	Coiffard, Mohr & Bernardes-de-Oliveira	Early Cretaceous	Crato Formation	Brazil	Fossil Aroid.
Trapa pannonica[39]	Sp. nov.	Valid	Hably	Late Miocene		Hungary	A water caltrop.
Trapa rozsaszentmartoni[39]	Sp. nov.	Valid	Hably	Late Miocene		Hungary	A water caltrop.
Virola dominicana[46]	Sp. nov.	Valid	Poinar & Steeves	Cenozoic (Eocene or Miocene)		Dominican Republic	A member of Myristicaceae found in Dominican amber, a species of Virola.
Vitis szakmanygyorgyi[39]	Sp. nov.	Valid	Hably	Late Miocene		Hungary	A species of Vitis.

Other seed plants

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Donponoxylon[47]	Gen. et 2 sp. nov.	Valid	Tidwell, Britt & Wright	Middle to Late Jurassic		Australia  New Zealand	A seed plant of uncertain phylogenetic placement. The type specimen is Donponoxylon bennettii; genus also contains Donponoxylon jacksonii.
Dordrechtites arcanus[48]	Sp. nov.	Valid	Bergene, Taylor & Taylor	Middle Triassic (Anisian)	Fremouw Formation	Antarctica	A gymnosperm of uncertain phylogenetic placement.

Other plants

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Acinosporites tristratus[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Jubah Formation	Saudi Arabia	A trilete spore.
Alatisporites? trisacculus[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Jauf Formation	Saudi Arabia	A trilete spore.
Biornatispora elegantula[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Jauf Formation	Belgium  Germany  Saudi Arabia	A trilete spore.
Biornatispora microclavata[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Jauf Formation	Saudi Arabia	A trilete spore.
Brochotriletes crameri[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Jauf Formation Jubah Formation	Saudi Arabia	A trilete spore.
Brochotriletes tenellus[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Jauf Formation	Saudi Arabia	A trilete spore.
Brochotriletes tripapillatus[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Awaynat Wanin II Formation	Libya	A trilete spore.
Camarozonotriletes asperulus[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Awaynat Wanin I Formation Awaynat Wanin II Formation Awaynat Wanin III Formation	Brazil  Tunisia	A trilete spore.
Chelinospora carnosa[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Jauf Formation	Saudi Arabia	A trilete spore.
Chelinospora condensata[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Jauf Formation	Saudi Arabia	A trilete spore.
Chelinospora densa[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Jauf Formation	Saudi Arabia	A trilete spore.
Chelinospora laxa[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Jauf Formation	Saudi Arabia	A trilete spore.
Chelinospora vulgata[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Jauf Formation	Saudi Arabia	A trilete spore.
Coronaspora inornata[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Jauf Formation	Saudi Arabia	A trilete spore.
Cristatisporites streelii[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Awaynat Wanin II Formation	Brazil  Tunisia	A trilete spore.
Cymbosporites variabilis[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Jauf Formation	Saudi Arabia	A trilete spore.
Cymbosporites variegatus[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Awaynat Wanin II Formation	Brazil  Libya  Saudi Arabia  Tunisia  United Kingdom	A trilete spore.
Cymbosporites wellmanii[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Jauf Formation	Saudi Arabia	A trilete spore.
Cyrtospora tumida[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Awaynat Wanin I Formation Awaynat Wanin II Formation Awaynat Wanin III Formation	Saudi Arabia  Tunisia	A trilete spore.
Diaphanospora milleri[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Jauf Formation Ouan Kasa Formation	Saudi Arabia	A trilete spore.
Dibolisporites tuberculatus[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Awaynat Wanin I Formation Awaynat Wanin II Formation Jauf Formation Jubah Formation	Brazil  Libya  Saudi Arabia  Spain[50]  Tunisia  United States	A trilete spore.
Dibolisporites verecundus[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Jauf Formation	Saudi Arabia	A trilete spore.
Dictyotriletes hemeri[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Jubah Formation	Saudi Arabia  Spain	A trilete spore.
Dictyotriletes marshallii[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Jauf Formation Ouan Kasa Formation	Saudi Arabia	A trilete spore.
Elenisporis gondwanensis[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Awaynat Wanin I Formation Awaynat Wanin II Formation Jubah Formation	Saudi Arabia  Tunisia	A trilete spore.
Emphanisporites laticostatus[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Awaynat Wanin III Formation Jubah Formation	Brazil  Saudi Arabia  Tunisia	A trilete spore.
Emphanisporites plicatus[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Jauf Formation Jubah Formation	Saudi Arabia	A trilete spore.
Franhueberia[51]	Gen. et sp. nov.	Valid	Hoffman & Tomescu	Devonian (Emsian)	Battery Point Formation	Canada	A basal member of Euphyllophytina. The type specimen is F. gerriennei.
Geminospora convoluta[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Awaynat Wanin II Formation Jubah Formation	Saudi Arabia	A trilete spore.
Grandispora maura[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Awaynat Wanin I Formation Awaynat Wanin II Formation Awaynat Wanin III Formation	Saudi Arabia  Tunisia	A trilete spore.
Granulatisporites concavus[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Awaynat Wanin I Formation Awaynat Wanin II Formation Jauf Formation Jubah Formation	Libya  Saudi Arabia  Spain[50]	A trilete spore.
Hystricosporites brevispinus[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Awaynat Wanin II Formation	Tunisia	A trilete spore.
Raistrickia commutata[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Awaynat Wanin II Formation Awaynat Wanin III Formation	Saudi Arabia  Tunisia	A trilete spore.
Raistrickia jaufensis[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Jauf Formation	Saudi Arabia	A trilete spore.
Retusotriletes atratus[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Jauf Formation Jubah Formation	Australia  Saudi Arabia  Spain[50]	A trilete spore.
Retusotriletes celatus[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Jauf Formation	Saudi Arabia	A trilete spore.
Samarisporites tunisiensis[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Ouan Kasa Formation	Tunisia	A trilete spore.
Verrucosisporites nafudensis[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Jauf Formation	Saudi Arabia	A trilete spore.
Verrucosisporites onustus[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Jauf Formation	Saudi Arabia	A trilete spore.
Verrucosisporites stictus[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Jauf Formation Ouan Kasa Formation	Saudi Arabia	A trilete spore.
Zonotriletes brevivelatus[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Awaynat Wanin I Formation Jauf Formation Jubah Formation Ouan Kasa Formation	Algeria  Libya  Saudi Arabia  Tunisia	A trilete spore.
Zonotriletes rotundus[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Awaynat Wanin I Formation Awaynat Wanin II Formation Ouan Kasa Formation	Brazil  Tunisia	A trilete spore.
Zonotriletes venatus[49]	Sp. nov	Valid	Breuer & Steemans	Devonian	Jauf Formation Ouan Kasa Formation	Algeria  Brazil  Saudi Arabia  Tunisia	A trilete spore.