2020 in arthropod paleontology

2020 in arthropod paleontology is a list of new arthropod fossil taxa, including arachnids, crustaceans, insects, trilobites, and other arthropods that were announced or described, as well as other significant arthropod paleontological discoveries and events which occurred in 2020.

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Arachnids

Newly named taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Aliendiguetia[1]	Gen. et sp. nov	Valid	Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider of uncertain phylogenetic placement, possibly related to the Ochyroceratoidea. The type species is A. praecursor.
Bicornoculus granulans[1]	Sp. nov	Valid	Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the family Tetrablemmidae.
Biungulus[2]	Gen. et sp. nov	In press	Bartel et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Opiliones belonging to the family Epedanidae. Genus includes new species B. xiai.
Burmacharon[3]	Gen. et sp. nov	In press	Hu et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A whip spider. Genus includes new species B. dunlopi.
Burmesarchaea bilongapophyses[1]	Sp. nov	Valid	Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the family Archaeidae.
Burmesia[1]	Gen. et sp. nov	Junior homonym	Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to family Nemesiidae. The type species is B. sordida. The generic name is preoccupied by Burmesia Healey (1908); Wunderlich (2021) coined a replacement name Burmesiana.[4]
?Burmesiola kachinensis[1]	Sp. nov	Valid	Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A tree trunk spider.
Burmorchestina circular[1]	Sp. nov	Valid	Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the family Oonopidae.
Burmorchestina prominens[1]	Sp. nov	Valid	Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the family Oonopidae.
Burmorsolus globosus[1]	Sp. nov	Valid	Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider, probably related to trogloraptorids.
Burmorsolus longembolus[1]	Sp. nov	Valid	Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider, probably related to trogloraptorids.
Burmorsolus longibulbus[1]	Sp. nov	Valid	Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider, probably related to trogloraptorids.
Chaeriloiurus[5]	Gen. et sp. nov	Disputed	Lourenço in Lourenço & Velten	Cretaceous	Burmese amber	Myanmar	A scorpion belonging to the family Chaerilobuthidae or Pseudochactidae.[6] Genus includes new species C. brigittemuellerae. Xuan et al. (2025) considered the genus Chaeriloiurus to be a junior synonym of the genus Chaerilobuthus, though the authors maintained C. brigittemuellerae as a distinct species within the latter genus.[6]
Chaeriloscorpiops[5]	Gen. et sp. nov	Valid	Lourenço in Lourenço & Velten	Cretaceous	Burmese amber	Myanmar	A scorpion belonging to the family Palaeoeuscorpiidae. Genus includes new species C. bautschi.
Consteniusi[7]	Gen. et sp. nov	Valid	Downen & Selden	Eocene	Kishenehn Formation	United States	An orb-weaver spider. The type species is C. leonae.
Curculioides[8]	Gen. et sp. nov	Valid	Whalen & Selden	Carboniferous (Pennsylvanian)	Energy Shale	United States	A member of Ricinulei. Genus includes new species C. bohemondi.
Cymbioblemma fusca[1]	Sp. nov	Valid	Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the family Tetrablemmidae.
Cymbioblemma hamoembolus[1]	Sp. nov	Valid	Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the family Tetrablemmidae.
Electroblemma bifurcata[1]	Sp. nov	Valid	Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the family Tetrablemmidae.
Electroblemma caula[1]	Sp. nov	Valid	Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the family Tetrablemmidae.
Electroblemma pinnae[1]	Sp. nov	Valid	Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the family Tetrablemmidae.
Ellenbergellus[2]	Gen. et sp. nov	In press	Bartel et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Opiliones belonging to the family Tithaeidae. Genus includes new species E. tuberculatus.
Eogamasomorpha rostratis[1]	Sp. nov	Valid	Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the family Tetrablemmidae.
Epieremulus sidorchukae[9]	Sp. nov	In press	Arillo, Subías & Peñalver	Cretaceous	Asturian amber	Spain	A mite belonging to the group Oribatida and to the family Caleremaeidae
Gigantocheles[2]	Gen. et sp. nov	In press	Bartel et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Opiliones belonging to the family Epedanidae. Genus includes new species G. nilsi.
Gigarachne[10]	Gen. et sp. nov	Valid	Jiang & Li in Jiang et al.	Late Cretaceous	Burmese amber	Myanmar	A spider belonging to the group Araneomorphae and to the new family Gigarachnidae. The type species is G. bian.
Greenwaltarachne[7]	Gen. et sp. nov	Valid	Downen & Selden	Eocene	Kishenehn Formation	United States	An orb-weaver spider. The type species is G. pamelae.
Hamoderces[1]	Gen. et sp. nov	Valid	Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the superfamily Scytodoidea and the family Praepholcidae. The type species is H. opilionoides.
Magnosegestria[1]	Gen. et sp. nov	Valid	Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	Probably a tube-dwelling spider. The type species is M. tuber.
Mesokanus[2]	Gen. et sp. nov	In press	Bartel et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Opiliones belonging to the group Laniatores and to the new family Mesokanidae. Genus includes new species M. oehmkuehnlei.
Mesozomus[11]	Gen. et sp. nov	Valid	Müller et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Schizomida. Genus includes new species M. groehni.
Modisimus chiapanecus[12]	Sp. nov	In press	García-Villafuerte & Valdez-Mondragón	Miocene (Aquitanian)	Mexican amber	Mexico	A species of Modisimus
Myannemesia[1]	Gen. et sp. nov	Valid	Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to family Nemesiidae. The type species is M. glaber.
Myrmecarchaea antecessor[13]	Sp. nov		Carbuccia et al.	Eocene (Ypresian)	Oise amber	France	A member of the family Archaeidae.
Neophyllobius electrus[14]	Sp. nov	Valid	Zmudzinski	Eocene	Baltic amber	Europe (Baltic Sea region)	A mite, a species of Neophyllobius.
Neophyllobius glaesus[14]	Sp. nov	Valid	Zmudzinski	Eocene	Baltic amber	Europe (Baltic Sea region)	A mite, a species of Neophyllobius.
Odontomegops[15]	Gen. et sp. nov	Valid	Guo et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the family Lagonomegopidae. Genus includes new species O. titan.
Odontoscirus cretacico[16]	Sp. nov	Valid	Porta et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A mite belonging to the family Bdellidae.
Palaeobeloniscus[2]	Gen. et sp. nov	In press	Bartel et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Opiliones belonging to the family Beloniscidae. Genus includes new species P. thilolebi.
Palazarqaraneus[1]	Gen. et sp. nov	Valid	Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to family Zarqaraneidae. The type species is P. hamulus.
Parvocteniza[1]	Gen. et sp. nov	Valid	Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to family Ctenizidae. The type species is P. parvula.
Paurospina fastigata[1]	Sp. nov	Valid	Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to family Zarqaraneidae.
Pilosarachne[10]	Gen. et sp. nov	Valid	Jiang & Li in Jiang et al.	Late Cretaceous	Burmese amber	Myanmar	A spider belonging to the group Araneomorphae and to the new family Pilosarachnidae. The type species is P. ju.
Praearaneus araneoides[1]	Sp. nov	Valid	Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the family Praearaneidae.
Priscaleclercera furcata[1]	Sp. nov	Valid	Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the family Psilodercidae.
Priscaleclercera hamo[1]	Sp. nov	Valid	Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the family Psilodercidae.
Priscaleclercera liber[1]	Sp. nov	Valid	Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the family Psilodercidae.
Procheiridium[17]	Gen. et sp. nov	Valid	Porta et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A pseudoscorpion belonging to the family Cheiridiidae and the subfamily Pycnocheiridiinae. Genus includes new species P. judsoni.
Propterpsiloderces crassitibia[1]	Sp. nov	Valid	Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the family Eopsilodercidae.
Propterpsiloderces cymbioseta[1]	Sp. nov	Valid	Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the family Eopsilodercidae.
Propterpsiloderces duplex[1]	Sp. nov	Valid	Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the family Eopsilodercidae.
Protertheraphosa[1]	Gen. et sp. nov	Valid	Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A tarantula. The type species is P. spinipes.
Protopyramidops[2]	Gen. et sp. nov	In press	Bartel et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Opiliones, possibly belonging to the family Pyramidopidae. Genus includes new species P. nalae.
Spinicreber vacuus[1]	Sp. nov	Valid	Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the family Pholcochyroceridae.
Suraju[18]	Gen. et sp. nov	In press	Martine et al.	Permian (Guadalupian)	Teresina Formation	Brazil	A scorpion. Genus includes new species S. itayma.
Unicornutiblemma[1]	Gen. et 3 sp. et comb. nov	Valid	Wunderlich & Müller	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to the family Tetrablemmidae. The type species is U. gracilicornis; genus also includes new species U. brevicornis and U. longicornis, as well as "Eogamasomorpha" unicornis Wunderlich (2017).

Crustaceans

Newly named taxa

Malacostracans

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Aliaplax[19]	Gen. et sp. nov	Valid	Pasini et al.	Early Pleistocene		Italy	A member of the family Goneplacidae. The type species is A. tyrsenorum.
Ammopylocheles romankijoki[20]	Sp. nov	Valid	Fraaije et al.	Early Cretaceous (Berriasian)		Czech Republic	A hermit crab belonging to the family Pylochelidae.
Amphoranina[21]	Gen. et 2 sp. nov	Valid	Nyborg et al.	Eocene to Oligocene		Canada United States	A crab belonging to the family Raninidae. Genus includes new species A. blandi and A. multispinata.
Ancipitecancer[22]	Gen. et sp. nov	Valid	Pasini, Luque & Garassino	Pleistocene		Italy	A member of Xanthoidea sensu lato, of uncertain phylogenetic placement within that group. Genus includes new species A. collinsi.
Angusteryon[23]	Gen. et sp. nov	Valid	Audo & Furrer	Early Jurassic		Switzerland	A member of Polychelida, possibly belonging to the family Coleiidae. Genus includes new species A. oberlii.
Annuntidiogenes sagittula[20]	Sp. nov	Valid	Fraaije et al.	Early Cretaceous (Berriasian)		Czech Republic	A hermit crab belonging to the family Annuntidiogenidae.
Bournelyreidus shiranui[24]	Sp. nov	Valid	Ando in Ando et al.	Late Cretaceous (Cenomanian)	Enokuchi Formation	Japan	A lyreidid raninoid crab.
Braggitrapezia[25]	Gen. et sp. nov	Valid	De Angeli & Caporiondo	Eocene		Italy	A crab belonging to the group Trapezioidea. Genus includes new species B. alessandroi.
Caecorchestia[26]	Gen. et sp. nov	Valid	Hegna & Lazo-Wasem in Hegna et al.	Early Miocene	Mexican amber	Mexico	A member of Amphipoda belonging to the family Talitridae. Genus includes new species C. bousfieldi.
Calappilia erwinhartei[27]	Sp. nov	Valid	Wallaard et al.	Miocene (Burdigalian)		Portugal	A species of Calappilia.
Callianassa reinhardpalorum[28]	Sp. nov	Valid	Hyžný	Miocene		Hungary Poland Slovakia	A species of Callianassa.
Calliax nishiki[29]	Sp. nov	Valid	Ando et al.	Latest Pleistocene to Holocene	Moeshima Silt Bed	Japan	A member of Axiidea belonging to the family Eucalliacidae.
Callichirus santensis[30]	Sp. nov	In press	Hernáez, Buchmann & Santana	Late Pleistocene		Brazil	A species of Callichirus.
Cancer nancei[31]	Sp. nov	Valid	Bahman, Feldmann & Schweitzer	Miocene	St. Marys	United States	A species of Cancer
Cancer zahrae[31]	Sp. nov	Valid	Bahman, Feldmann & Schweitzer	Miocene	St. Marys	United States	A species of Cancer
Cherpihomola[32]	Gen. et sp. nov	Valid	Marangon & De Angeli	Oligocene (Rupelian)		Italy	A member of the family Homolidae. The type species is C. italica.
Collinscarcinus[33]	Gen. et sp. nov	Valid	Artal & van Bakel	Eocene (Ypresian)		Spain	A crab belonging to the family Xanthidae. Genus includes new species C. obliquesulcatus.
Cretacalliax[34]	Gen. et sp. nov	Valid	Pasini, Poore & Garassino	Late Cretaceous (Cenomanian)		Lebanon	A ghost shrimp belonging to the family Callianopsidae. Genus includes new species C. levantina.
Cuchiadromites[35]	Gen. et sp. nov	In press	Ossó, van Bakel & Ferratges	Early Cretaceous (Aptian)	Patrocinio Formation	Spain	A crab belonging to the family Longodromitidae. Genus includes new species C. jadeae.
Daira corallina[36]	Sp. nov	Valid	Ferratges, Zamora & Aurell	Late Eocene		Spain
Dardanus cherpionensis[37]	Sp. nov	Valid	Marangon & De Angeli	Oligocene (Rupelian)	Ligure-Piemontese Basin	Italy	A species of Dardanus.
Dardanus faxensis[38]	Sp. nov	Valid	Jakobsen et al.	Paleocene (Danian)	Faxe Formation	Denmark	A species of Dardanus.
Diogenes augustinus[39]	Sp. nov	Valid	Ferratges, Zamora & Aurell	Eocene (Bartonian)	Arguis Formation	Spain	A hermit crab, a species of Diogenes.
Discosalaputium[40]	Gen. et sp. nov	Valid	Schädel et al.	Late Triassic (Carnian)	Reingraben	Austria	A member of Isopoda belonging to the group Scutocoxifera. The type species is D. aschauerorum.
Dynomene collinsi[41]	Sp. nov	Valid	Ossó et al.	Eocene (Priabonian)		Spain	A crab belonging to the family Dynomenidae.
Eodromites bernchrisdomiorum[42]	Sp. nov	Valid	Klompmaker et al.	Late Jurassic (Oxfordian)		Poland	A crab belonging to the family Goniodromitidae.
Eodromites cristinarobinsae[42]	Sp. nov	Valid	Klompmaker et al.	Early Cretaceous (Albian)	Eguino Formation	Spain	A crab belonging to the family Goniodromitidae.
Eopolycheles[43]	Gen. et sp. nov	Valid	Gašparič et al.	Late Triassic (Carnian)		Slovenia	A member of the family Polychelidae. Genus includes new species E. cornuaureus.
Eryma moriedaorum[24]	Sp. nov	Valid	Ando et al.	Late Cretaceous (Cenomanian)	Enokuchi Formation	Japan	A member of the family Erymidae.
Etyus tresgalloi[44]	Sp. nov	Valid	Ossó & Moreno-Bedmar	Early Cretaceous (Aptian)		Spain	A crab belonging to the family Etyiidae.
Eupilumnus? vokesae[45]	Sp. nov	In press	Luque in Luque et al.	Miocene		Mexico	A crab.
Europrosopon[42]	Gen. et comb. nov	Valid	Klompmaker et al.	Late Jurassic		Austria Czech Republic Germany Romania	A crab belonging to the family Prosopidae. The type species is "Prosopon" aculeatum von Meyer (1857); genus also includes "Prosopon" abbreviatum Schweitzer & Feldmann (2009).
Galathea tuscia[19]	Sp. nov	Valid	Pasini et al.	Early Pleistocene		Italy	A species of Galathea.
Gammaroidorum[46]	Gen. et sp. nov	In press	Jarzembowski et al.	Early Cretaceous (Hauterivian)	Weald	United Kingdom	A member of Amphipoda. Genus includes new species G. vonki.
Gelrincola[40]	Gen. et sp. nov	Valid	Schädel et al.	Middle Triassic (Anisian)	Lower Muschelkalk Vossenveld Formation	Netherlands	A member of Isopoda belonging to the group Cymothoida. The type species is G. winterswijkensis.
Gemmellarocarcinus riglosensis[36]	Sp. nov	Valid	Ferratges, Zamora & Aurell	Late Eocene		Spain
Glebocarcinus doii[47]	Sp. nov	Valid	Karasawa & Takahashi	Early Miocene	Igami	Japan	A member of the family Cancridae
Graptocarcinus collinsi[48]	Sp. nov	Valid	Vega in Vega et al.	Late Cretaceous (Maastrichtian)	Arroyo Grande Formation	Cuba	A crab belonging to the family Dynomenidae.
Hefriga schlechtingerae[49]	Sp. nov	Valid	Winkler	Late Jurassic (Tithonian)	Solnhofen	Germany	A caridean shrimp
Hispanigalathea raymondcaseyi[50]	Sp. nov	Valid	Van Bakel et al.	Early Cretaceous (Albian)	Gault	United Kingdom	A member of Galatheoidea.
Hoploparia echinata[51]	Sp. nov	Valid	Pinheiro et al.	Late Cretaceous (Santonian–Campanian)	Santa Marta Formation	Antarctica
Hoploparia hinokunika[24]	Sp. nov	Valid	Ando & Shimada in Ando et al.	Late Cretaceous (Cenomanian)	Enokuchi Formation Karakizaki Formation	Japan	A lobster.
Hoploparia tectumque[52]	Sp. nov	In press	Fraaije et al.	Late Cretaceous (Cenomanian)		Kazakhstan
Icriobranchiocarcinus rodas[53]	Sp. nov	In press	Vega in Vega et al.	Late Cretaceous (Maastrichtian)	Arroyo Grande Formation	Cuba	A crab belonging to group Portunoidea and the family Lithophylacidae. Announced in 2020; the final version of the article naming it is scheduled to be published in 2021.
Joecalcinus[54]	Gen. et sp. nov	Valid	Fraaije et al.	Eocene		Italy	A hermit crab belonging to the family Calcinidae. Genus includes new species J. bolcensis.
Juranecrocarcinus[55]	Gen. et comb. nov	In press	Van Bakel et al.	Late Jurassic (Kimmeridgian)		Germany	A crab belonging to the family Necrocarcinidae. The type species is "Nodoprosopon" angulosum Wehner (1988).
Laeviprosopon musialiki[56]	Sp. nov	Valid	Starzyk	Late Jurassic (Oxfordian)		Poland	A crab belonging to the family Prosopidae.
Levashidromites[57]	Gen. et sp. nov	In press	Van Bakel et al.	Early Cretaceous (Aptian)	Kubina Formation	Russia	A crab belonging to the family Longodromitidae. The type species is L. cornutus.
Lobogalenopsis joei[36]	Sp. nov	Valid	Ferratges, Zamora & Aurell	Late Eocene		Spain
Mariaplax ohiranorikoae [58]	Sp. nov	Valid	Karasawa and Kobayashi	Pleistocene	Atsumi Group	Japan	A member of the family Hexapodidae.
Marmacuma[59]	Gen. et sp. nov	Valid	Artüz & Sakınç	Late Pleistocene		Turkey	A member of Cumacea belonging to the family Nannastacidae. Genus includes new species M. samimei.
Masticacheles septemgradu[20]	Sp. nov	Valid	Fraaije et al.	Late Jurassic (Tithonian)		Czech Republic	A hermit crab belonging to the family Pilgrimchelidae.
Matutites collinsi[60]	Sp. nov	Valid	Davis, Garassino & Weaver	Middle Eocene	Castle Hayne	United States	A member of the family Matutidae.
Maurocarpilius[61]	Gen. et sp. nov	Valid	Ossó, Gagnaison & Bailleul	Eocene (Ypresian)	Ait Ouarhitane	Morocco	A member of the family Carpiliidae. The type species is M. binodosus.
Mesogalathea ardua[62]	Sp. nov	Valid	Gašparič, Robins & Gale	Late Jurassic (Kimmeridgian/Tithonian)		Slovenia	A member of the family Galatheidae.
Mesoparapylocheles janetjacksonae[20]	Sp. nov	Valid	Fraaije et al.	Late Jurassic (Tithonian)		Czech Republic	A hermit crab belonging to the family Parapylochelidae.
Mizuhocancer[47]	Gen. et comb. nov	Valid	Karasawa in Karasawa & Takahashi	Early—Middle Miocene		Japan	A member of the family Cancridae. Monotypic with the type species, Cancer? Imamurae Imaizumi, 1962.
Montemagrecarcinus[63]	Gen. et sp. nov	Valid	De Angeli & Ceccon	Early Eocene		Italy	A crab belonging to the family Pilumnidae. Genus includes new species M. efremi.
Moravacarcinus[55]	Gen. et sp. nov	In press	Van Bakel et al.	Early Cretaceous (Berriasian)	Štramberk Limestone	Czech Republic	A crab belonging to the family Necrocarcinidae. The type species is M. stramberkensis.
Munidopsis kaedetatsuyai [64]	Sp. nov	Valid	Karasawa	Miocene	Higashibessho Formation	Japan	A member of the family Munidopsidae.
Neoliomera zovoensis[65]	Sp. nov	Valid	De Angeli & Garassino	Early Eocene		Italy	A species of Neoliomera.
Nitotacarcinus ambrosii[66]	Sp. nov	Valid	Beschin & Checchi	Eocene (Ypresian)		Italy	A crab belonging to the superfamily Carpilioidea and the family Tumidocarcinidae.
Obtusotelson[40]	Gen. et sp. nov	Valid	Schädel et al.	Late Triassic (Carnian)	Reingraben	Austria	A member of Isopoda belonging to the group Scutocoxifera. The type species is O. summesbergeri.
Orithyia eikii[67]	Sp. nov	Valid	Karasawa	Early Miocene	Mizunami Group	Japan	A member of the family Orithyiidae.
Paguristes collinsi[37]	Sp. nov	Valid	Marangon & De Angeli	Oligocene (Rupelian)	Ligure-Piemontese Basin	Italy	A species of Paguristes.
Paguristes frigoscopulus[38]	Sp. nov	Valid	Jakobsen et al.	Paleocene (Danian)	Faxe Formation	Denmark	A species of Paguristes.
Paguristes joecollinsi[68]	Sp. nov	Valid	Wallaard et al.	Miocene (Tortonian)	Pakhna Formation	Cyprus	A species of Paguristes.
Paguristes laurae[37]	Sp. nov	Valid	Marangon & De Angeli	Oligocene (Rupelian)	Ligure-Piemontese Basin	Italy	A species of Paguristes.
Palaeopalinurellus[69]	Gen. et 2 sp. et comb. nov	Valid	Fraaije et al.	Late Jurassic to Late Cretaceous (Cenomanian)[70]		Czech Republic France Lebanon[70]	A furry lobster. Genus includes new species P. culocervus and P. jbeilensis,[70] as well as "Palinurus" strambergensis Bachmayer (1959).
Palaeopolycheles nantosueltae[71]	Sp. nov		Jauvion et al.	Middle Jurassic (Callovian)	La Voulte-sur-Rhône	France	A member of Polychelida
Personadorippe[57]	Gen. et 2 sp. et comb. nov	In press	Van Bakel et al.	Cretaceous (Aptian to Cenomanian)	Hudspeth Formation Kubina Formation Lyaminsk Formation	Russia United States	A crab belonging to the superfamily Dorippoidea and the family Telamonocarcinidae. The type species is P. kalashnikovi; genus also includes new species P. levashiensis, as well as "Eodorippe" connori Nyborg et al. (2019).
Petrolisthes landsendi[72]	Sp. nov	Valid	Garassino & Nyborg in Garassino et al.	Late Cretaceous (Santonian)	Nanaimo Group	Canada	A species of Petrolisthes.
?Proeryon erinaceus[73]	Sp. nov	Valid	Audo, Schweigert & Charbonnier	Late Jurassic (Oxfordian)	Terrain à Chailles Formation	France
?Proeryon quiltyi[73]	Sp. nov	Valid	Audo, Schweigert & Charbonnier	Middle Jurassic (Callovian)		Antarctica
Prosopon josephcollinsi[42]	Sp. nov	Valid	Klompmaker et al.	Late Jurassic (Tithonian)	Ernstbrunn Limestone	Austria	A crab belonging to the family Prosopidae.
Protoapseudoidus[74]	Fam. et Gen. et sp. nov	In press	Heard et al.	Early Cretaceous (Aptian)	Sierra Madre	Mexico	A member of Tanaidacea belonging to the group Apseudomorpha, in the new family Protoapseudoidae. Genus includes new species P. espinalensis.
Protopagurus cerebellum[20]	Sp. nov	Valid	Fraaije et al.	Early Cretaceous (Berriasian)		Czech Republic	A hermit crab belonging to the family Paguridae.
Protopagurus duopupae[20]	Sp. nov	Valid	Fraaije et al.	Early Cretaceous (Berriasian)		Czech Republic	A hermit crab belonging to the family Paguridae.
Pseudastacus lemovices[75]	Sp. nov	Valid	Charbonnier & Audo	Early Jurassic (Sinemurian)		France	A member of Astacidea belonging to the family Stenochiridae
Sabellidromites lanae[76]	Sp. nov	Valid	Nyborg, Garassino & Ross	Late Cretaceous (late Campanian)		Canada	A dromioid crab
Stenodactylina beardi[77]	Sp. nov	Valid	Feldmann, Schweitzer & Haggart	Late Cretaceous	Haslam	Canada	A member of the family Erymidae.
Stenodactylina devillezi[78]	Sp. nov	Valid	Schweigert & Härer	Late Jurassic (Kimmeridgian)	Nusplingen	Germany	A member of the family Erymidae.
Stenodactylina geigerae[78]	Sp. nov	Valid	Schweigert & Härer	Late Jurassic (Tithonian)	Usseltal	Germany	A member of the family Erymidae.
Stonesfielderyon[73]	Gen. et sp. nov	Valid	Audo, Schweigert & Charbonnier	Middle Jurassic (Bathonian)		United Kingdom	A possible member of the family Eryonidae. The type species is S. imitator.
Tanidromites nightwishorum[42]	Sp. nov	Valid	Klompmaker et al.	Late Jurassic (Tithonian)	Ernstbrunn Limestone	Austria	A crab belonging to the family Tanidromitidae.
Tanidromites weinschenki[42]	Sp. nov	Valid	Klompmaker et al.	Late Jurassic (Kimmeridgian)	Oberjura-Massenkalk Formation	Germany	A crab belonging to the family Tanidromitidae.
Tymolus bottemilleri[79]	Sp. nov	Valid	Nyborg & Garassino	Miocene	Astoria	United States	A crab belonging to the family Cyclodorippidae.
Tymolus collinsi[79]	Sp. nov	Valid	Nyborg & Garassino	Miocene		United States	A crab belonging to the family Cyclodorippidae.
Upogebia azorensis[80]	Sp. nov	Valid	Hyžný in Hyžný et al.	Pliocene		Azores	A species of Upogebia. Announced in 2020; the final version of the article naming it was published in 2021.
Vecticallichirus kawanoi[81]	Sp. nov	Valid	Ando	Late Eocene	Funazu	Japan	A ghost shrimp belonging to the family Callichiridae.
Vectis collinsi[57]	Sp. nov	In press	Van Bakel et al.	Early Cretaceous (Aptian)	Kubina Formation	Russia	A crab belonging to the family Viaiidae.

Ostracods

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Adelphobolbina praeclara[82]	Sp. nov	Valid	Becker, Braun & Pratt	Devonian		Canada
Ampuloides beckeri[83]	Sp. nov	Valid	Nazik et al.	Late Devonian		Mongolia	A member of Podocopida belonging to the superfamily Bairdiocypridoidea and the family Pachydomellidae.
Angarallina[84]	Gen. et 2 sp. nov	Valid	Melnikova	Ordovician		Russia	Genus includes new species A. demissa and A. aenigma.
Araxobairdia[85]	Gen. et sp. nov	Valid	Gliwa in Gliwa et al.	Permian (Changhsingian)		Iran	A member of Podocopida belonging to the family Bairdiidae. The type species is A. formosa.
Bairdia andrecrasquini[86]	Sp. nov		Crasquin et al.	Late Triassic		Italy	A member of the family Bairdiidae.
Bairdia gambaneraensis[86]	Sp. nov		Crasquin et al.	Late Triassic		Italy	A member of the family Bairdiidae.
Bairdia paramolesta[87]	Sp. nov	Valid	Cabral, Lord & Pinto in Cabral et al.	Early Jurassic (Pliensbachian and Toarcian)		Portugal	A member of the family Bairdiidae.
Bairdiacypris argonautaii[88]	Sp. nov	Valid	Forel in Forel & Grădinaru	Late Triassic (Rhaetian)	Kössen Formation	Austria Romania	A member of Podocopida belonging to the family Bairdiidae.
Bairdiacypris kathleenae[85]	Sp. nov	Valid	Gliwa in Gliwa et al.	Late Permian and Early Triassic (Changhsingian–Griesbachian)		Iran	A member of Podocopida belonging to the family Bairdiidae.
Beyrichiopsis hushootensis[83]	Sp. nov	Valid	Nazik et al.	Late Devonian		Mongolia	A member of Platycopida belonging to the superfamily Kloedenelloidea and the family Beyrichiopsidae.
Bythocypris dorsogibba[89]	Sp. nov	In press	Trabelsi et al.	Early Cretaceous (Albian)	Orbata	Tunisia
Bythocypris? multagracilis[89]	Sp. nov	In press	Trabelsi et al.	Early Cretaceous (Albian)	Orbata	Tunisia
Bythocypris? pythagorasi[89]	Sp. nov	In press	Trabelsi et al.	Early Cretaceous (Aptian)	Orbata	Tunisia
Candonopsis carthaginensis[89]	Sp. nov	In press	Trabelsi et al.	Early Cretaceous (Albian)	Orbata	Tunisia
Capsacythere baldanzae[90]	Sp. nov	In press	Sciuto & Temani in Sciuto, Temani & Ammar	Miocene (Messinian)		Tunisia	A member of Podocopa belonging to the family Trachyleberididae.
Capsacythere fekihi[90]	Sp. nov	In press	Sciuto & Temani in Sciuto, Temani & Ammar	Miocene (Messinian)		Tunisia	A member of Podocopa belonging to the family Trachyleberididae.
Capsacythere giovannae[90]	Sp. nov	In press	Sciuto & Temani in Sciuto, Temani & Ammar	Miocene (Messinian)		Tunisia	A member of Podocopa belonging to the family Trachyleberididae.
Capsacythere salaji[90]	Sp. nov	In press	Sciuto & Temani in Sciuto, Temani & Ammar	Miocene (Messinian)		Tunisia	A member of Podocopa belonging to the family Trachyleberididae.
Carinaknightina hofmanni[85]	Sp. nov	Valid	Gliwa in Gliwa et al.	Permian (Changhsingian)		Iran	A member of Palaeocopida belonging to the family Kirkbyidae.
Cavellina fosteri[85]	Sp. nov	Valid	Gliwa in Gliwa et al.	Permian (Changhsingian)		Iran	A member of Platycopida belonging to the family Cavellinidae.
Cavellina hairapetiani[85]	Sp. nov	Valid	Gliwa in Gliwa et al.	Permian (Changhsingian)		Iran	A member of Platycopida belonging to the family Cavellinidae.
Ceratobairdia? akhilleus[88]	Sp. nov	Valid	Forel in Forel & Grădinaru	Late Triassic (Rhaetian)		Romania	A member of Podocopida belonging to the family Bairdiidae.
Chrysocythere arutai[90]	Sp. nov	In press	Sciuto & Temani in Sciuto, Temani & Ammar	Miocene (Messinian)		Tunisia	A member of Podocopa belonging to the family Trachyleberididae.
Chrysocythere gliozziae[90]	Sp. nov	In press	Sciuto & Temani in Sciuto, Temani & Ammar	Miocene (Messinian)		Tunisia	A member of Podocopa belonging to the family Trachyleberididae.
Chrysocythere russoi[90]	Sp. nov	In press	Sciuto & Temani in Sciuto, Temani & Ammar	Miocene (Messinian)		Tunisia	A member of Podocopa belonging to the family Trachyleberididae.
Cimbaurila maamourii[90]	Sp. nov	In press	Sciuto & Temani in Sciuto, Temani & Ammar	Miocene (Messinian)		Tunisia	A member of Podocopa belonging to the family Hemicytheridae.
Costa ponticulocarinata[91]	Sp. nov	Valid	Yasuhara et al.	Early Miocene		India
Costoprimites multicostatus[92]	Sp. nov	Valid	Gonta	Ordovician		Russia
Costoprimites processus[92]	Sp. nov	Valid	Gonta	Ordovician		Russia
Cypridea punctacentralis[93]	Sp. nov	In press	Santos Filho, Fauth & Sames	Early Cretaceous	Orós Formation	Brazil
Cytherella ilariae[90]	Sp. nov	In press	Sciuto & Temani in Sciuto, Temani & Ammar	Miocene (Messinian)		Tunisia	A species of Cytherella.
Cytheretta mariaantoniettae[90]	Sp. nov	In press	Sciuto & Temani in Sciuto, Temani & Ammar	Miocene (Messinian)		Tunisia	A member of Podocopa belonging to the family Cytherettidae.
Dicerobairdia buseri[94]	Sp. nov	Valid	Forel in Forel, Kolar-Jurkovšek & Jurkovšek	Late Triassic (Carnian)		Slovenia	A member of Podocopida belonging to the family Bairdiidae.
Direigina[84]	Gen. et sp. nov	Valid	Melnikova	Ordovician		Russia	Genus includes new species D. nana.
Disulcina proxima[84]	Sp. nov	Valid	Melnikova	Ordovician		Russia	A member of the family Tetradellidae.
Dolocythere amphistiela[95]	Sp. nov	Valid	Lord, Cabral & Danielopol	Early Cretaceous (Albian)		Germany	A member of Podocopida belonging to the family Cytheridae
Dolocytheridea donzei[89]	Sp. nov	In press	Trabelsi et al.	Early Cretaceous (Albian)	Orbata	Tunisia
Dorsogibbella alium[92]	Sp. nov	Valid	Gonta	Ordovician		Russia
Dorukella crasquinae[90]	Sp. nov	In press	Sciuto & Temani in Sciuto, Temani & Ammar	Miocene (Messinian)		Tunisia	A member of Podocopa belonging to the family Hemicytheridae.
Dorukella nevioi[90]	Sp. nov	In press	Sciuto & Temani in Sciuto, Temani & Ammar	Miocene (Messinian)		Tunisia	A member of Podocopa belonging to the family Hemicytheridae.
Dorukella razgallahae[90]	Sp. nov	In press	Sciuto & Temani in Sciuto, Temani & Ammar	Miocene (Messinian)		Tunisia	A member of Podocopa belonging to the family Hemicytheridae.
Dorukella ruggeroi[90]	Sp. nov	In press	Sciuto & Temani in Sciuto, Temani & Ammar	Miocene (Messinian)		Tunisia	A member of Podocopa belonging to the family Hemicytheridae.
Eumiraculum mettei[85]	Sp. nov	Valid	Gliwa in Gliwa et al.	Permian (Wuchiapingian–Changhsingian)		Iran	A member of Podocopida belonging to the superfamily Bairdioidea.
Fabalicypris veronicae[85]	Sp. nov	Valid	Gliwa in Gliwa et al.	Permian (Changhsingian)		Iran	A member of Podocopida belonging to the family Bairdiidae.
Foramenella sibirica[84]	Sp. nov	Valid	Melnikova	Ordovician		Russia
Graptocythere octopus[90]	Sp. nov	In press	Sciuto & Temani in Sciuto, Temani & Ammar	Miocene (Messinian)		Tunisia	A member of Podocopa belonging to the family Hemicytheridae.
Hallatina oblonga[84]	Sp. nov	Valid	Melnikova	Ordovician		Russia	A member of the group Leperditellocopida belonging to the family Primitiidae.
Hastacypris[93]	Gen. et sp. nov	Junior homonym	Santos Filho, Fauth & Sames	Early Cretaceous	Orós Formation	Brazil	A member of Cypridoidea. Genus includes new species H. adamantem. The generic name is preoccupied by Hastacypris Croneis & Gutke (1939); Guillam (2024) coined a replacement name Cuspicypris.[96]
Heterocypris aspirasensis[97]	Sp. nov	Valid	Durak, Akkiraz & Nazik	Early Miocene	Aspiras Basin	Turkey	A species of Heterocypris. Announced in 2020; the final version of the article naming it was published in 2021.
Histriabairdia[88]	Gen. et sp. et comb. nov	Valid	Forel in Forel & Grădinaru	Late Triassic (Carnian) to Middle Jurassic (Bathonian)		Austria France Germany India Italy Romania  Switzerland	A member of Podocopida belonging to the family Bairdiidae. The type species is H. pontuseuxinusensis; genus also includes "Anchistrocheles"? spinosa Sheppard in Brand (1990), "Anchistrocheles"? tuningensis Beher (2004) and "Anchistrocheles" gemmellaroi Crasquin in Crasquin et al. (2018).
Hungarella forelae[86]	Sp. nov		Crasquin et al.	Late Triassic		Italy	A member of the family Healdiidae.
Hungarella siciliiensis[86]	Sp. nov		Crasquin et al.	Late Triassic		Italy	A member of the family Healdiidae.
Hungaroleberis striatus[85]	Sp. nov	Valid	Gliwa in Gliwa et al.	Permian (Changhsingian)		Iran	A member of Myodocopida belonging to the family Cylindroleberididae.
Illativella triquetra[82]	Sp. nov	Valid	Becker, Braun & Pratt	Devonian		Canada
Ilyocypris? arca[93]	Sp. nov	In press	Santos Filho, Fauth & Sames	Early Cretaceous	Orós Formation	Brazil
Ilyocypris? latanodi[93]	Sp. nov	In press	Santos Filho, Fauth & Sames	Early Cretaceous	Orós Formation	Brazil
Isobythocypris atalantella[88]	Sp. nov	Valid	Forel in Forel & Grădinaru	Late Triassic (Rhaetian)		Romania	A member of Podocopida belonging to the family Bairdiidae.
Jonesites chunensis[92]	Sp. nov	Valid	Gonta	Ordovician		Russia
Kerocythere dittainoensis[86]	Sp. nov		Crasquin et al.	Late Triassic		Italy	A member of the family Cytheruridae.
Kiesowia (?) incerta[84]	Sp. nov	Valid	Melnikova	Ordovician		Russia	A member of the family Ctenonotellidae.
Kinkelinella ventrocarinata[87]	Sp. nov	Valid	Cabral, Pinto & Lord in Cabral et al.	Early Jurassic (Toarcian)		Portugal United Kingdom	A member of the family Protocytheridae.
Kinnekullea gaia[98]	Sp. nov	Valid	Hearing et al.	Ordovician (Katian)	Phu Ngu	Vietnam
Korolyukina[84]	Gen. et 2 sp. nov	Valid	Melnikova	Ordovician		Russia	A member of the family Bolliidae. Genus includes new species K. conspicua and K. taimyrica.
Laccochilina (?) jugata[84]	Sp. nov	Valid	Melnikova	Ordovician		Russia	A member of the family Eurychilinidae.
Laccochilina (Prochilina) yanpingensis[99]	Sp. nov	In press	Zhang	Ordovician	Shihtzupu Formation	China
Leucocytherella dangeloi[100]	Sp. nov	Valid	Alivernini in Alivernini et al.	Miocene (Messinian) and Pliocene	Zhada Basin	China	A member of the family Limnocytheridae.
Liuzhinia julfensis[85]	Sp. nov	Valid	Gliwa in Gliwa et al.	Late Permian and Early Triassic (Changhsingian–Griesbachian)		Iran	A member of Podocopida belonging to the superfamily Bairdioidea.
Lyumellina[92]	Gen. et sp. nov	Valid	Gonta	Ordovician		Russia	Genus includes new species L. risus
Minyocythere[95]	Gen. et 2 sp. et comb. nov	Valid	Lord, Cabral & Danielopol	Middle Jurassic (Aalenian and Bajocian)		Germany Netherlands United Kingdom	A cytherid Podocopida. The type species is M. macroporosa; genus also includes new species M. angulata, as well as "Dolocythere" maculosa Bate (1963) and "Dolocythere" tuberculata Luppold (2012).
Mockella barbroae[86]	Sp. nov		Crasquin et al.	Late Triassic		Italy	A member of the family Limnocytheridae.
Mongolianella kyranbeki[101]	Sp. nov	In press	Choi et al.	Cretaceous	Yanji	China
Myanmarcypris[102]	Gen. et sp. nov	Valid	Wang et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Candonidae belonging to the subfamily Paracypridinae. Genus includes new species M. hui.
Nikolina entonipteros[92]	Sp. nov	Valid	Gonta	Ordovician		Russia
Nikolina pteroventralis[92]	Sp. nov	Valid	Gonta	Ordovician		Russia
Okadaleberis (?) benzartiae[90]	Sp. nov	In press	Sciuto & Temani in Sciuto, Temani & Ammar	Miocene (Messinian)		Tunisia	A member of Podocopa belonging to the family Trachyleberididae.
Okadaleberis decimai[90]	Sp. nov	In press	Sciuto & Temani in Sciuto, Temani & Ammar	Miocene (Messinian)		Tunisia	A member of Podocopa belonging to the family Trachyleberididae.
Okadaleberis guerneti[90]	Sp. nov	In press	Sciuto & Temani in Sciuto, Temani & Ammar	Miocene (Messinian)		Tunisia	A member of Podocopa belonging to the family Trachyleberididae.
Okadaleberis hendae[90]	Sp. nov	In press	Sciuto & Temani in Sciuto, Temani & Ammar	Miocene (Messinian)		Tunisia	A member of Podocopa belonging to the family Trachyleberididae.
Okadaleberis khayatii[90]	Sp. nov	In press	Sciuto & Temani in Sciuto, Temani & Ammar	Miocene (Messinian)		Tunisia	A member of Podocopa belonging to the family Trachyleberididae.
Okadaleberis memmiae[90]	Sp. nov	In press	Sciuto & Temani in Sciuto, Temani & Ammar	Miocene (Messinian)		Tunisia	A member of Podocopa belonging to the family Trachyleberididae.
Orthobairdia capuliformis[85]	Sp. nov	Valid	Gliwa in Gliwa et al.	Permian (Changhsingian)		Iran	A member of Podocopida belonging to the family Bairdiidae.
Paijenborchellina reitanoi[90]	Sp. nov	In press	Sciuto & Temani in Sciuto, Temani & Ammar	Miocene (Messinian)		Tunisia	A member of Podocopa belonging to the family Cytheridae.
Paractinocythereis[91]	Gen. et comb. nov	Valid	Yasuhara et al.	Early Miocene		India	A new genus for "Actinocythereis" gujaratensis Tewari & Tandon (1960)
Paracypris chekhmai[89]	Sp. nov	In press	Trabelsi et al.	Early Cretaceous (Aptian)	Orbata	Tunisia
Paracypris ovidi[88]	Sp. nov	Valid	Forel in Forel & Grădinaru	Late Triassic (Rhaetian)		Romania	A member of Cypridoidea belonging to the family Paracyprididae.
Perissocytheridea altesulcata[89]	Sp. nov	In press	Trabelsi et al.	Early Cretaceous (Aptian)	Orbata	Tunisia
Perissocytheridea kouminiensis[89]	Sp. nov	In press	Trabelsi et al.	Early Cretaceous (Albian)	Orbata	Tunisia
Petasobairdia amazonella[88]	Sp. nov	Valid	Forel in Forel & Grădinaru	Late Triassic (Rhaetian)		Romania	A member of the family Bairdiidae.
Petasobairdia jeandercourti[86]	Sp. nov		Crasquin et al.	Late Triassic		Italy	A member of the family Bairdiidae.
Pontocypris? karavankensis[94]	Sp. nov	Valid	Forel in Forel, Kolar-Jurkovšek & Jurkovšek	Late Triassic (Carnian)		Slovenia	A member of Cypridoidea belonging to the family Pontocyprididae.
Postsinusella[92]	Gen. et sp. nov	Valid	Gonta	Ordovician		Russia	Genus includes new species P. guttaformis.
Prasuchonella? huarpe[103]	Sp. nov	Valid	Carignano, Echevarría & Zavattieri	Middle Triassic	Cuyo Basin	Argentina	A member of Darwinulocopina.
Pseudoleperditia? vetusta[82]	Sp. nov	Valid	Becker, Braun & Pratt	Devonian		Canada
Pseudomacrocypris? kerabani[88]	Sp. nov	Valid	Forel in Forel & Grădinaru	Late Triassic (Rhaetian)		Romania	A member of Cypridoidea belonging to the family Pontocyprididae.
Pseudulrichia replicata[84]	Sp. nov	Valid	Melnikova	Ordovician		Russia	A member of the family Richinidae.
Ptychobairdia iudicaensis[86]	Sp. nov		Crasquin et al.	Late Triassic		Italy	A member of the family Bairdiidae.
Ptychobairdia leonardoi[86]	Sp. nov		Crasquin et al.	Late Triassic		Italy	A member of the family Bairdiidae.
Pythagoracypris[93]	Gen. et sp. nov	In press	Santos Filho, Fauth & Sames	Early Cretaceous	Orós Formation	Brazil	Genus includes new species P. latavectis.
Rhinocypris? ericius[93]	Sp. nov	In press	Santos Filho, Fauth & Sames	Early Cretaceous	Orós Formation	Brazil
Ruggieria quadricarinata[90]	Sp. nov	In press	Sciuto & Temani in Sciuto, Temani & Ammar	Miocene (Messinian)		Tunisia	A member of Podocopa belonging to the family Cytherettidae.
Steusloffia rudiformis[92]	Sp. nov	Valid	Gonta	Ordovician		Russia
Sulcicuneus gemmifer[82]	Sp. nov	Valid	Becker, Braun & Pratt	Devonian		Canada
Tasmanocypris lochardi[104]	Sp. nov	In press	Warne	Late Miocene and Pliocene		Australia
Tesakovites[84]	Gen. et sp. nov	Valid	Melnikova	Ordovician		Russia	A member of the family Circulinidae. Genus includes new species T. rotundatus.
Urftella triangula[82]	Sp. nov	Valid	Becker, Braun & Pratt	Devonian		Canada
Urocythere (Pokornyella) salaktaensis[90]	Sp. nov	In press	Sciuto & Temani in Sciuto, Temani & Ammar	Miocene (Messinian)		Tunisia	A member of Podocopa belonging to the family Hemicytheridae.
Velatomorpha rochacamposi[105]	Sp. nov	Valid	Bergue, Maranhao & Ng	Permian		Brazil

Other crustaceans

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Actinobalanus? sloveniensis[106]	Sp. nov	Valid	Buckeridge et al.	Oligocene and Miocene		Slovenia	A barnacle.
Altiverruca capsa[107]	Sp. nov	Valid	Gale	Pliocene-early Pleistocene		Indian Ocean (Mascarene Plateau)	A barnacle belonging to the family Verrucidae.
Altiverruca fusione[107]	Sp. nov	Valid	Gale	Pliocene-early Pleistocene		Indian Ocean (Mascarene Plateau)	A barnacle belonging to the family Verrucidae.
Ambocyclus[108]	Gen. et comb. nov	Valid	Schweitzer, Mychko & Feldmann	Carboniferous		Ireland Russia United States?	A member of Cyclida. Genus includes A. capidulum (Chernyschev, 1933), A. simulans (Reed, 1908) and possibly A. ? minutus (Rogers, 1902).
Angulotergum kiamichensis[109]	Sp. nov	Valid	Gale	Cretaceous		United States	A barnacle.
Angulotergum milviformis[109]	Sp. nov	Valid	Gale	Cretaceous		United States	A barnacle.
Angulotergum robustum[109]	Sp. nov	Valid	Gale	Cretaceous		United States	A barnacle.
Arcoscalpellum s.l. isaonishikawai [110]	Sp. nov	Valid	Karasawa	Miocene	Katsuta Group	Japan	A member of the family Scalpellidae.
Arcoscalpellum s.l. Joei [110]	Sp. nov	Valid	Karasawa	Miocene	Yotsuyaku Formation	Japan	A member of the family Scalpellidae.
Arcuatoscalpellum wacoensis[109]	Sp. nov	Valid	Gale	Cretaceous		United States	A barnacle.
Ashinkailepas indica[111]	Sp. nov	Valid	Gale in Gale et al.	Late Pleistocene		India	A barnacle
Aurivillialepas mascarensis[112]	Sp. nov	Valid	Gale	Pleistocene		Indian Ocean (Mascarene Plateau)	A barnacle belonging to the family Calanticidae.
Brachylepas hantonensis[113]	Sp. nov	Valid	Gale	Cretaceous		United Kingdom	A barnacle
Brachylepas thieli[113]	Sp. nov	Valid	Gale	Cretaceous		Germany	A barnacle
Brittaniclus[108]	Gen. et comb. nov	Valid	Schweitzer, Mychko & Feldmann	Carboniferous		United Kingdom	A member of Cyclida belonging to the family Americlidae. Genus includes B. rankini (Woodward, 1868), B. scotti (Woodward, 1894) and B. testudo (Peach, 1882).
Carabicyclus[108]	Gen. et comb. nov	Valid	Schweitzer, Mychko & Feldmann	Carboniferous		Ireland United Kingdom	A member of Cyclida. Genus includes C. wrighti (Woodward, 1870).
Catherinum busselli[107]	Sp. nov	Valid	Gale	Pliocene-early Pleistocene		Indian Ocean (Mascarene Plateau)	A barnacle belonging to the family Scalpellidae.
Chernyshevine[108]	Gen. et comb. nov	Valid	Schweitzer, Mychko & Feldmann	Carboniferous		Tajikistan	A member of Cyclida. Genus includes C. spinosus (Chernyschev, 1933).
Collinslepas[113]	Gen. et 3 sp. nov	Valid	Gale	Cretaceous		Morocco Tunisia	A barnacle. Genus includes new species C. aitlaminensis, C. robustus and C. tunisiensis.
Costatolepas[107]	Gen. et sp. nov	Valid	Gale	Pliocene-early Pleistocene		Indian Ocean (Mascarene Plateau)	A barnacle belonging to the family Scalpellidae. The type species is C. buckeridgei.
Costatoverruca baxteri[107]	Sp. nov	Valid	Gale	Pliocene-early Pleistocene		Indian Ocean (Mascarene Plateau)	A barnacle belonging to the family Verrucidae.
Costatoverruca macropluteum[112]	Sp. nov	Valid	Gale	Pleistocene		Indian Ocean (Mascarene Plateau)	A barnacle belonging to the family Verrucidae.
Costatoverruca tredecima[112]	Sp. nov	Valid	Gale	Pleistocene		Indian Ocean (Mascarene Plateau)	A barnacle belonging to the family Verrucidae.
Cretiscalpellum robaszynskii[114]	Sp. nov	In press	Gale, Jagt & Goolaerts	Late Cretaceous (Maastrichtian)		Tunisia	A barnacle.
Daohugounaias[115]	Gen. et sp. nov	Valid	Luo et al.	Middle Jurassic	Daohugou	China	A member of Anostraca. Genus includes new species D. cheni.
Diotascalpellum acies[109]	Sp. nov	Valid	Gale	Cretaceous		United States	A barnacle.
Dziklus[108]	Gen. et comb. nov	Valid	Schweitzer, Mychko & Feldmann	Carboniferous	Mazon Creek fossil beds	United States	A member of Cyclida belonging to the family Americlidae. Genus includes D. obesus (Schram, Vonk & Hof, 1997).
Eoverruca aubensis[113]	Sp. nov	Valid	Gale	Early Cretaceous (Albian)		France	A barnacle
Eoverruca symmetrica[113]	Sp. nov	Valid	Gale	Late Cretaceous (Campanian)		United Kingdom	A barnacle
Etcheslepas portlandensis[116]	Sp. nov	Valid	Gale	Tithonian–Berriasian	Portland Group	United Kingdom	A barnacle
Fallaxlepas[113]	Gen. et comb. nov	Valid	Gale	Cretaceous		United Kingdom	A brachylepadid barnacle. The type species is "Pollicipes" fallax Darwin (1851)
Gibbosaverruca youngi[107]	Sp. nov	Valid	Gale	Pliocene-early Pleistocene		Indian Ocean (Mascarene Plateau)	A barnacle belonging to the family Verrucidae.
Iliestheria urumqiensis[117]	Sp. nov	Valid	Teng & Li	Early Jurassic (Pliensbachian–Toarcian)	Sangonghe	China	A clam shrimp. Announced in 2020; the final version of the article naming was published in 2021.
Ivolepas worthensis[109]	Sp. nov	Valid	Gale	Cretaceous		United States	A barnacle.
Jagtscalpellum[113]	Gen. et comb. et 3 sp. nov	Valid	Gale	Cretaceous		Tunisia United Kingdom United States	A barnacle. The type species is "Pollicipes" striatus Darwin (1851); genus also includes "Pollicipes" filosus Withers (1911), as well as new species J. africanum, J. spinosum and J. luteorum.[109]
Lahuerguinagrapta[118]	Gen. et sp. nov	In press	Gallego et al.	Early Cretaceous	La Huérguina	Spain	An afrograptid clam shrimp. Genus includes new species L. multicostata.
Litocyclus[108]	Gen. et comb. nov	Valid	Schweitzer, Mychko & Feldmann	Carboniferous		Ireland United Kingdom United States?	A member of Cyclida. Genus includes L. bilobatus (Woodward, 1870), L. jonesianus (Woodward, 1870), L. torosus (Woodward, 1870) and possibly also L. ? communis (Rogers, 1902) and L. ? permarginatus (Rogers, 1902).
Lophobalanus fresvillensis[119]	Sp. nov	Valid	Gale	Eocene (Lutetian)		France	A barnacle. Announced in 2020; the final version of the article naming it was published in 2021.
Loriculina fragila[109]	Sp. nov	Valid	Gale	Cretaceous		United States	A barnacle.
Loriculina rubrifluvia[109]	Sp. nov	Valid	Gale	Cretaceous		United States	A barnacle.
Loriolepas whytei[116]	Sp. nov	Valid	Gale	Tithonian–Berriasian		United Kingdom	A barnacle
Newmaniverruca multitabulata[107]	Sp. nov	Valid	Gale	Pliocene-early Pleistocene		Indian Ocean (Mascarene Plateau)	A barnacle belonging to the family Verrucidae.
Pachylasma traceyi[112]	Sp. nov	Valid	Gale	Pliocene-Pleistocene		Indian Ocean (Mascarene Plateau)	A barnacle belonging to the family Pachylasmatidae.
Proverruca anglica[113]	Sp. nov	Valid	Gale	Cretaceous		United Kingdom	A barnacle
Regioscalpellum kennedyi[113]	Sp. nov	Valid	Gale	Cretaceous		Tunisia	A barnacle
Rostratoverruca darwini[107]	Sp. nov	Valid	Gale	Pliocene-early Pleistocene		Indian Ocean (Mascarene Plateau)	A barnacle belonging to the family Verrucidae.
Striascalpellum[113]	Gen. et 3 sp. et comb. nov	Valid	Gale	Cretaceous		Sweden United Kingdom	A barnacle. The type species is S. elegans; genus also includes new species S. barringtonensis (subsequently transferred to the genus Virgilepas)[120] and S. bromleyi, as well as "Scalpellum (Titanolepas)" martini Withers (1935).
Subsecolepas[113]	Gen. et sp. nov	Valid	Gale	Cretaceous		United Kingdom	A barnacle. The type species is S. holtwilsoni.
Tazawacyclus[108]	Gen. et comb. nov	Valid	Schweitzer, Mychko & Feldmann	Carboniferous		Japan	A member of Cyclida. Genus includes T. tazawai (Niko & Ibaraki, 2011).
Texaslepas[109]	Gen. et sp. nov	Valid	Gale	Cretaceous		United States	A barnacle. Genus includes new species T. holterhoffi.
Vectibalanus[119]	Gen. et comb. et sp. nov	Valid	Gale	Eocene and Oligocene	Barton Bouldnor Headon Hill Selsey	United Kingdom	A barnacle. The type species is "Balanus" unguiformis Sowerby (1846); genus also includes "Balanus" erisma Sowerby (1846) and a new species V. mortoni. Announced in 2020; the final version of the article naming it was published in 2021.
Virgilepas[113]	Gen. et 3 sp. nov	Valid	Gale	Cretaceous		Morocco United Kingdom	A barnacle. Genus includes new species V. aboudaensis, V. peakei and V. hancocki.
Virgiscalpellum laevis[113]	Sp. nov	Valid	Gale	Cretaceous		United Kingdom	A barnacle. Subsequently made the type species of the separate genus Laeviscalpellum.[120]
Virgiscalpellum marysensis[109]	Sp. nov	Valid	Gale	Cretaceous		United States	A barnacle.
Virgiscalpellum mhrilensis[113]	Sp. nov	Valid	Gale	Cretaceous		Tunisia	A barnacle
Virgiscalpellum multilineatum[113]	Sp. nov	Valid	Gale	Cretaceous		Tunisia	A barnacle
Virgiscalpellum rugosum[109]	Sp. nov	Valid	Gale	Cretaceous		United States	A barnacle. Subsequently transferred to the genus Laeviscalpellum.[120]
Virgiscalpellum sussexiense[113]	Sp. nov	Valid	Gale	Cretaceous		United Kingdom	A barnacle
Virgiscalpellum truncatum[113]	Sp. nov	Valid	Gale	Cretaceous		United Kingdom	A barnacle
Witherscalpellum[113]	Gen. et comb. nov	Valid	Gale	Cretaceous		France United Kingdom	A barnacle. The type species is "Cretiscalpellum aptiensis Withers (1935); genus also includes "Cretiscalpellum" matrioni Gale (2019).

General research

• Redescription and a study on the phylogenetic relationships of Francocaris grimmi, based on data from new fossil material from the Solnhofen Limestone (Germany), is published by Pazinato et al. (2020).^[121]
• A hermit crab belonging to the family Pylochelidae preserved in a shell of an ammonite belonging to the species Craspedites nekrassovi will be described from the Upper Jurassic of Moscow, Russia by Mironenko (2020).^[122]
• A putative lobster Tricarina gadvanensis from the Early Cretaceous of Iran is reinterpreted as a seroloid isopod by Hyžný et al. (2020).^[123]
• A study on the phylogenetic relationships of Eocarcinus praecursor is published by Scholtz (2020).^[124]
• A study on anatomical adaptations that enabled an ecological transition into the water column in Silurian myodocopes, focusing on myodocopids from the Herefordshire Lagerstätte (United Kingdom), is published by Siveter, Perrier & Williams (2020).^[125]
• A study comparing sexual dimorphism in ostracods from the Gulf and Atlantic coastal plain before and after the Cretaceous–Paleogene extinction event is published by Martins et al. (2020).^[126]
• Redescription of Americlus rankini and a study on the paleoecology of this taxon is published by Clark et al. (2020).^[127]

Insects

Main article: 2020 in insect paleontology

Trilobites

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Acastoides biebrichensis[128]	Sp. nov	Valid	Alberti	Devonian (Emsian)	Rupbach Shale	Germany	A member of the family Acastidae.
Acastoides requadti[128]	Sp. nov	Valid	Alberti	Devonian (Emsian)	Rupbach Shale	Germany	A member of the family Acastidae.
Aciculolenus askewi[129]	Sp. nov	Valid	Chatterton	Cambrian (Jiangshanian)		Canada
Arduennella hainauensis[130]	Sp. nov	Valid	Van Viersen & Alberti	Devonian (Emsian)		Germany
Atlanticalymene[131]	Gen. et sp. nov	Valid	Adrain, Karim & McAdams	Ordovician (Darriwilian)	Table Cove Formation	Canada	A calymenine trilobite. The type species is A. bardensis.
Balvibole[132]	Gen. et sp. nov	Valid	Basse & Lemke	Devonian (Famennian)	Dasberg Limestone	Germany	A cyrtosymboline trilobite. Genus includes new species B. kaufmanni.
Blountia angelae[133]	Sp. nov	Valid	Armstrong, Westrop & Eoff	Cambrian	Nolichucky Formation	United States	A member of the family Kingstoniidae.
Blountia morgancreekensis[133]	Sp. nov	Valid	Armstrong, Westrop & Eoff	Cambrian	Riley Formation	United States	A member of the family Kingstoniidae.
Blountia nevadensis[133]	Sp. nov	Valid	Armstrong, Westrop & Eoff	Cambrian	Mendha Formation	United States	A member of the family Kingstoniidae.
Blountia newfoundlandensis[133]	Sp. nov	Valid	Armstrong, Westrop & Eoff	Cambrian	Shallow Bay Formation	Canada	A member of the family Kingstoniidae.
Blountia tennesseensis[133]	Sp. nov	Valid	Armstrong, Westrop & Eoff	Cambrian	Nolichucky Formation	United States	A member of the family Kingstoniidae.
Boethiusia[134]	Gen. et comb. nov	Valid	Geyer	Cambrian		Italy	A new genus for "Dolerolenus" courtessolei Pillola (1991).
Clappaspis tupeq[135]	Sp. nov	Valid	Peel	Cambrian (Miaolingian)	Telt Bugt Formation	Greenland	A member of the family Alokistocaridae.
Cliffia nicoleae[129]	Sp. nov	Valid	Chatterton	Cambrian (Jiangshanian)		Canada
Cyphaspides paulospinatus[136]	Sp. nov	Valid	Alberti	Devonian		Germany	A member of Proetida belonging to the family Aulacopleuridae.
Cyphaspides speculator[136]	Sp. nov	Valid	Alberti	Devonian		Germany	A member of Proetida belonging to the family Aulacopleuridae.
Dicranurus webbyi[137]	Sp. nov	Valid	Holloway, Smith & Thomas	Ordovician (Katian)	Gunningbland Formation	Australia	An odontopleurid trilobite.
Ehmaniella sermersuaqensis[135]	Sp. nov	Valid	Peel	Cambrian (Miaolingian)	Telt Bugt Formation	Greenland	A member of the family Alokistocaridae.
Elrathia hensonensis[138]	Nom. nov	Valid	Geyer & Peel	Cambrian	Ekspedition Bræ Formation	Greenland	A replacement name for Elrathia groenlandica Geyer & Peel (2017).
Eowinterbergia effenbergensis[132]	Sp. nov	Valid	Basse & Lemke	Devonian (Famennian)	Wocklum Limestone	Germany
Epicurella[134]	Gen. et sp. nov	Valid	Geyer	Cambrian		Morocco	A member of the family Despujolsiidae. Genus includes new species E. pustulata.
Feistops[139]	Gen. et 2 sp. nov	Valid	Crônier et al.	Devonian (Famennian)		Mongolia	A phacopid trilobite. Genus includes new species F. mongoliensis and F. khovdensis. Announced in 2020; the final version of the article naming was published in 2021.
Gravicalymene bakeri[140]	Sp. nov	In press	Smith & Ebach	Ordovician (Katian)	Gordon Group	Australia	A calymenid trilobite.
?Houseops olonbulagensis[139]	Sp. nov	Valid	Crônier et al.	Devonian (Famennian)		Mongolia	A phacopid trilobite. Announced in 2020; the final version of the article naming was published in 2021.
Kingaspidoides spinirecurvatus[141]	Sp. nov	Valid	Geyer, Pais & Wotte	Cambrian	Jbel Wawrmast Formation	Morocco	A member of Redlichiida belonging to the family Ellipsocephalidae.
Marsaisia devoillei[134]	Sp. nov	Valid	Geyer	Cambrian		Morocco	A member of the family Despujolsiidae.
Marsaisia? lemdadensis[134]	Sp. nov	Valid	Geyer	Cambrian		Morocco	A member of the family Despujolsiidae.
Phantaspis[142]	Gen. et sp. nov	Valid	Sun, Zeng & Zhao	Cambrian (Wuliuan)	Mantou Formation	China	A member of Ptychopariacea of uncertain phylogenetic placement. The type species is P. auritus.
Proabadiella[143]	Gen. et 2 sp. nov	Valid	Liñán et al.	Cambrian Stage 3	Pusa Formation	Spain	Genus includes new species P. toletana and P. vidalii.
Prophalaron[137]	Gen. et sp. nov	Valid	Holloway, Smith & Thomas	Ordovician (Katian)	Gunningbland Formation	Australia	A calymenid trilobite. Genus includes new species P. jonesi.
Prosaukia oculata[144]	Sp. nov	Valid	Wernette et al.	Cambrian (Furongian)	Ao Mo Lae Formation	Thailand
Satunarcus[145]	Gen. et sp. nov	Valid	Wernette et al.	Late Cambrian	Ao Mo Lae Formation	Thailand	A kaolishaniid trilobite belonging to the new subfamily Ceronocarinae. Genus includes new species S. molaensis.
Saukianda? dresnayi[146]	Sp. nov	In press	Geyer & Landing	Cambrian		Moroccan‒Algerian border region
Treveropyge hellemondi[147]	Sp. nov	Valid	Van Viersen & Taghon	Devonian (Emsian)	Our Formation	Belgium	A member of the family Acastidae.
Utia debra[148]	Sp. nov	Valid	Sundberg	Cambrian (Wuliuan)	Lakeview Limestone Spence Shale	United States	A member of Ptychopariida belonging to the family Utiidae.

General research

• A study on the timing of the trilobite extinctions at the early to middle Cambrian transition is published by Sundberg et al. (2020).^[149]
• Evidence of gregarious moulting of Cambrian (Stage 4) trilobites from the "Tsinghsutung" Formation (China), representing one of the earliest records of synchronized moulting behaviours in the fossil record, is presented by Corrales-García et al. (2020).^[150]
• A study on trilobite size variations in the Ordovician Fezouata Formation (Morocco) deposited in a cold-water environment is published by Saleh et al. (2020).^[151]
• A study on the nature of variation among oryctocephalid specimens from the Combined Metals Member of the Pioche Formation (Nevada; Cambrian Stage 4) will be published by Webster & Sundberg (2020).^[152]
• A study on the ontogeny of the olenid trilobite Leptoplastides salteri is published by Månsson & Clarkson (2020).^[153]
• A study on the anatomy of the internal structures of the compound eye of Aulacopleura koninckii is published by Schoenemann & Clarkson (2020).^[154]
• A study on exuviae of Ovalocephalus tetrasulcatus from the Ordovician Linhsiang Formation (Hubei, China) is published by Zong (2020), who reports exuviae arranged with two or three together end to end or superimposed, and evaluates the implications of these findings for the knowledge of the behavior of trilobites.^[155]

Other arthropods

Newly named taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Adelophthalmus pyrrhae[156]	Sp. nov	Valid	Lamsdell et al.	Carboniferous (Tournaisian)	St. Nazaire Group (possibly Lydiennes Formation)	France	A eurypterid.
Allolimulus[157]	Gen. et comb. nov	Disputed	Lamsdell	Jurassic		United Kingdom	A horseshoe crab. The type species is "Limulus" woodwardi Watson (1909). However, Bicknell et al. (2021) transferred "L." woodwardi to the genus Mesolimulus.[158]	(A) Norilimulus woodae, (B) Batracholimulus fuchsbergensis, (C) Boeotiaspis longispinus, (D) Keuperlimulis vicensis, (E) Volanalimulus madagascarensis, (F) Allolimulus woodwardi
Andersoniella[157]	Gen. et comb. nov	Valid	Lamsdell	Carboniferous		Germany United States	A member of the family Belinuridae. The type species is "Euproops" longispina Packard (1885).
Batracholimulus[157]	Gen. et comb. nov	Valid	Lamsdell	Triassic		Germany	A member of the family Austrolimulidae. The type species is "Paleolimulus" fuchsbergensis Hauschke & Wilde (1987).
Boeotiaspis[157]	Gen. et comb. nov	Valid	Lamsdell	Carboniferous		United States	A member of the family Austrolimulidae. The type species is "Paleolimulus" longispinus Schram (1979).
Bushizheia[159]	Gen. et sp. nov	Valid	O'Flynn & Liu in O'Flynn et al.	Cambrian Stage 3	Chiungchussu	China	An early euarthropod of uncertain phylogenetic placement. The type species is B. yangi.
Concavicaris martinae[160]	Sp. nov	Valid	Broda, Rak & Hegna	Devonian (Famennian)		Czech Republic	A thylacocephalan.
Concavicaris pikae[160]	Sp. nov	Valid	Broda, Rak & Hegna	Devonian (Famennian)		Poland	A thylacocephalan.
Concavicaris submarinus[161]	Sp. nov		Jobbins, Haug & Klug	Devonian (Famennian)		Morocco	A thylacocephalan.
Concavicaris woodruffensis[160]	Sp. nov	Valid	Broda, Rak & Hegna	Devonian (Famennian)	Woodruff Formation	United States	A thylacocephalan.
Devonopilio[162]	Gen. et sp. nov	Valid	Tihelka, Tian & Cai	Devonian (Pragian)	Rhynie chert	United Kingdom	Originally described as a member of Opiliones, but this interpretation was subsequently questioned by Pérez-González & Shultz (2021), who were unsure about the phylogenetic placement of this arthropod, but noted anatomical similarities with the euthycarcinoid species Heterocrania rhyniensis.[163] The type species is D. hutchinsoni.
Diplacanthocaris[164]	Gen. et sp. nov	Valid	Ji et al.	Early Triassic		China	A thylacocephalan. Genus includes new species D. chaohuensis.
Dolesea[165]	Gen. et sp. nov	Valid	Hannibal & May	Permian (Cisuralian)	Richards Spur locality	United States ( Oklahoma)	A millipede belonging to the group Helminthomorpha. The type species is D. subtila.
Ericixerxes[166]	Gen. et sp. nov		Gueriau et al.	Devonian (Famennian)	Evieux	Belgium	A member of Euthycarcinoidea. The type species is E. potii.
Hipponicharion perforata[167]	Sp. nov	Valid	Skovsted et al.	Cambrian	Mural	Canada	A member of Bradoriida belonging to the family Hipponicharionidae.
Itagnostus idahoensis[148]	Sp. nov	Valid	Sundberg	Cambrian (Wuliuan)	Emigrant Formation Lakeview Limestone	United States	A member of Agnostida belonging to the family Peronopsidae.
Kanoshoia[168]	Gen. et sp. nov	Valid	Lerosey-Aubril et al.	Cambrian (Drumian)	Wheeler Shale	United States	A megacheiran. Genus includes new species K. rectifrons.
Karstiulus[165]	Gen. et sp. nov	Valid	Hannibal & May	Permian (Cisuralian)	Richards Spur locality	United States ( Oklahoma)	A millipede belonging to the group Juliformia, possibly a member of the superfamily Xyloiuloidea. The type species is K. fortsillensis.
Keuperlimulus[157]	Gen. et comb. nov	Valid	Lamsdell	Triassic		France	A horseshoe crab. The type species is "Limulus" vicensis Bleicher (1897).
Macrobelinurus[157]	Gen. et comb. nov	Valid	Lamsdell	Carboniferous		United Kingdom	A member of the family Belinuridae. The type species is "Steropis" arcuatus Baily (1859).
Mesolimulus tafraoutensis[169]	Sp. nov	In press	Lamsdell et al.	Late Cretaceous (Cenomanian–Turonian)	Gara Sbaa Lagerstätte	Morocco	A horseshoe crab.
Microcaris rectilineatus[164]	Sp. nov	Valid	Ji et al.	Early Triassic		China	A thylacocephalan. Ehiro & Kano (2024) considered it to be a species belonging to the genus Parisicaris.[170]
Norilimulus[157]	Gen. et comb. nov	Valid	Lamsdell	Carboniferous (Tournaisian)	Horton Bluff Formation	Canada	A member of Xiphosura belonging to the family Paleolimulidae. The type species is "Paleolimulus" woodae Lerner, Lucas & Mansky (2016).
Oklahomasoma[165]	Gen. et sp. nov	Valid	Hannibal & May	Permian (Cisuralian)	Richards Spur locality	United States ( Oklahoma)	A millipede belonging to the group Helminthomorpha. The type species is O. richardsspurense.
Parabelinurus[157]	Gen. et comb. nov	Valid	Lamsdell	Carboniferous		Russia Ukraine United States	A member of the family Belinuridae. The type species is "Enthomolithus" lunatus Martin (1809); genus also includes P. iswariensis (Chernyshev, 1928), P. lacoei (Packard, 1885), P. metschetensis (Chernyshev, 1928) and P. stepanovi (Chernyshev, 1928).
Paraconcavicaris[160]	Gen. et comb. nov	Valid	Broda, Rak & Hegna	Carboniferous (Tournaisian)	Líšeň Formation	Czech Republic	A thylacocephalan; a new genus for "Concavicaris" viktoryni Rak, Broda & Kumpan (2018).
Parioscorpio[171]	Gen. et sp. nov		Wendruff et al.	Silurian (Telychian)	Brandon Bridge	United States	An arthropod of uncertain phylogenetic placement. Originally classified as an early scorpion, but subsequently argued to be basal to crown group Mandibulata and Chelicerata.[172] The type species is P. venator.
Patesia[173]	Gen. et comb. nov	Valid	Bicknell & Smith	Late Devonian		United States	A stem xiphosurid; a new genus for "Kasibelinurus" randalli. Announced in 2020; the final version of the article naming it was published in 2021.
Pauropsxenus extraneus[174]	Sp. nov	In press	Su, Cai & Huang	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A millipede belonging to the family Polyxenidae.
Pauropsxenus ordinatus[174]	Sp. nov	In press	Su, Cai & Huang	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A millipede belonging to the family Polyxenidae.
Pruemopterus[175]	Gen. et sp. nov	Valid	Poschmann	Early Devonian		Germany	A eurypterid. Genus includes new species P. salgadoi. Announced in 2020; the final version of the article naming it was published in 2021.
Pseudobeyrichona taurata[167]	Sp. nov	Valid	Skovsted et al.	Cambrian	Mural	Canada	A member of Bradoriida belonging to the family Hipponicharionidae.
Shpineviolimulus[176]	Gen. et comb. nov		Bicknell, Naugolnykh & Brougham	Permian (Asselian)	Araukaritovaya Formation	Ukraine	A member of Xiphosura belonging to the group Limulina; a new genus for "Paleolimulus" jakovlevi Glushenko & Ivanov (1961).
Sklerolibyon[177]	Gen. et sp. nov		Aria et al.	Cambrian Stage 3	Maotianshan Shale Member of the Yu'anshan Formation	China	A megacheiran related to Jianfengia and Fortiforceps. The type species is S. maomima.
Soligorskopterus shpinevi[178]	Sp. nov	Valid	Naugolnykh & Areshin	Late Devonian		Russia	An eurypterid. Announced in 2019; the final version of the article naming it was published in 2020.
Tariccoia tazagurtensis[179]	Sp. nov	Valid	Pérez-Peris et al.	Early Ordovician	Fezouata	Morocco	A nektaspid arthropod. Announced in 2020; the final version of the article naming it was published in 2021.
Thelxiope holmani[180]	Sp. nov	Valid	Lerosey-Aubril, Skabelund & Ortega-Hernández	Cambrian (Drumian)	Wheeler Shale	United States	A possible member of Chelicerata.
Volanalimulus[157]	Gen. et sp. nov	Valid	Lamsdell	Early Triassic	Ankitokazo Basin	Madagascar	A horseshoe crab. The type species is V. madagascarensis.
Xiaocaris[181]	Gen. et sp. nov		Liu et al.	Cambrian Stage 3	Chiungchussu Formation	China	A member of the group Fuxianhuiida. The type species is X. luoi.
Xiazhuangocaris[182]	Gen. et sp. nov	Valid	Zeng et al.	Cambrian Stage 3	Hongjingshao Formation	China	A member of Hymenocarina. Genus includes new species X. chenggongensis. Announced in 2020; the final version of the article naming it was published in 2021.

General research

• Exceptionally preserved specimens of Chuandianella ovata carrying a large number of small eggs are described from the Cambrian Chengjiang biota (China) by Ou et al. (2020), who compare the reproductive strategies of C. ovata and closely related Waptia fieldensis.^[183]
• Sun, Zeng & Zhao (2020) report the discovery of a member of the genus Sidneyia belonging or related to the species S. inexpectans from the Wuliuan Mantou Formation of North China, expanding known distribution of this genus.^[184]
• A putative xiphosuran Kiaeria is reinterpreted as a member of Chasmataspidida by Lamsdell (2020a).^[185]
• A study on the anatomy of head structures of Heterocrania rhyniensis, evaluating its implications for the knowledge of the phylogenetic relationships of euthycarcinoids, is published by Edgecombe et al. (2020).^[186]
• A study on heterochrony in Xiphosura, showing that clades that occupy non-marine environments show elevated rates of peramorphosis or paedomorphosis is published by Lamsdell (2020b).^[187]
• A study on the morphology of the feeding apparatuses of Eurypterida and Trigonotarbida is published by Haug (2020).^[188]
• A study aiming to determine the range of prey sizes that could be captured by sweep-feeding eurypterids is published by Hughes & Lamsdell (2020).^[189]
• A study on the anatomy and phylogenetic relationships of Anhelkocephalon handlirschi is published by Schädel & Haug (2020), who interpret this taxon as a probable member of Cyclida rather than an isopod.^[190]
• Liu et al. (2020) report evidence of the presence of a three-dimensionally preserved labrum associated with the mouth opening in juvenile specimens of Leanchoilia illecebrosa.^[191]