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Arrup
Genus of Mecistocephalidae centipedes From Wikipedia, the free encyclopedia
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Arrup is a genus of soil centipedes in the family Mecistocephalidae.[1] This genus contains sixteen species.[2] These centipedes are found mainly in temperate regions of East Asia with some species found in Central Asia and California.[3][4] Most species in this genus are soil-dwellers, but the Japanese species Arrup akiyoshiensis was discovered in a cave and might be a troglobiont.[5]
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Description
Centipedes in this genus range from 1 cm to 5 cm in length.[3] All species in this genus have 41 leg-bearing segments.[6][7] The body is homogeneous in pigmentation, without darker patches. The head in this genus features a transverse suture on the front of the dorsal surface. The side pieces of the labrum are fully divided into anterior and posterior sclerites.[8] The clypeus in this genus is almost completely areolate and features a longitudinal areolate stripe down the middle.[3][9] The pleurites on the side of the head lack setae. The coxosternite of the first maxillae is not divided down the middle by a longitudinal suture, and the coxosternite of the second maxillae is also undivided. The telopodites of the second maxillae are too short to reach beyond the first maxillae. The forcipular tergum is wider than long, with no longitudinal groove down the middle. The first article of the forcipule features one distal tooth, and the ultimate article features one basal tooth. The groove on the ventral surface of the trunk segments is not forked. The ultimate legs of the male are as slender as those of the female.[9][10]
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Phylogeny
A cladistic analysis of the family Mecistocephalidae using morphological features places the genus Arrup in the subfamily Arrupinae, along with the genera Agnostrup, Nannarrup, and Partygarrupius.[4] Further cladistic analysis of the subfamily Arrupinae based on external morphology places the genus Arrup in a clade with Nannarrup as a closely related sister group.[8] This analysis also places this clade inside another clade with Agnostrup as a sister group in the same branch of a phylogenetic tree.[8]
The genus Arrup shares some distinctive features with its close relatives in the genus Nannarrup. For example, in both genera, the setae on the clypeus are arranged in two groups, one on each side of the clypeus. Furthermore, like all Arrup species, all Nannarrup species have 41 pairs of legs. Other features, however, distinguish Arrup from Nannarrup. For example, the coxosternite of the first maxillae is divided in Nannarrup but undivided in Arrup.[8]
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Species
This genus currently includes sixteen accepted species:[1][2][11]
- Arrup akiyoshiensis Tsukamoto & Shimano, 2019
- Arrup areolatus (Shinohara, 1957)
- Arrup asiaticus (Titova, 1975)
- Arrup dentatus (Takakuwa, 1934)
- Arrup doii (Takakuwa, 1940)
- Arrup edentulus (Attems, 1904)
- Arrup holstii (Pocock, 1895)
- Arrup ishiianus Uliana, Bonato & Minelli, 2007
- Arrup kyushuensis Uliana, Bonato & Minelli, 2007
- Arrup lilliputianus Uliana, Bonato & Minelli, 2007
- Arrup longicalix Uliana, Bonato & Minelli, 2007
- Arrup mamaevi (Titova, 1975)
- Arrup obtusus (Takakuwa, 1934)
- Arrup pauroporus (Takakuwa, 1936)
- Arrup pylorus Chamberlin, 1912
- Arrup sauteri (Silvestri, 1919)
References
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