CRuMs

CRuMs is a clade of microbial eukaryotes, whose name is an acronym of the following constituent groups: i) Collodictyonids also known as diphylleids, ii) rigifilids and iii) mantamonads as sister of the Amorphea.^[4][1] A new CRuMs order Glissandrida was proposed in 2025 to place the yet unclassified genus Glissandra.^[5] It more or less supersedes Varisulca, as Ancyromonadida are inferred not to be specifically related to the orders Diphylleida/Collodictyonida, Rigifilida and Mantamonadida.

CRuMs
Collodictyon pseudopodoa
Scientific classification
Domain:	Eukaryota
Clade:	CRuMs Brown et al. 2018[1]
Subgroups
GRiDi Glissandrida Hilomonadea Diphyllatea Glissodiscea
Synonyms
Crumalia Zmitrovich, Perelygin & Zharikov 2022[2] Varisulca Cavalier-Smith 2013 emend. 2021[3]

Phylogeny

In 2025, GRiDi clade was proposed by Euki Yazaki et al.^[5]

Opimoda

Ancyromonadida Cavalier-Smith 1998 emend. Atkins 2000 Malawimonadea Cavalier-Smith 2003 Amorphea Amoebozoa Lühe 1913 emend. Cavalier-Smith 1998 Obazoa Breviatea Cavalier-Smith 2004 Apusomonadida Karpov & Mylnikov 1989 Opisthokonta (inc. fungi & animals) CRuMs Glissodiscea: Mantamonadida Cavalier-Smith 2004 GRiDi Glissandrida[5] Hilomonadea: Rigifilida Karpov & Mylnikov 1989 Diphyllatea: Diphylleida Cavalier-Smith 1993