Ceratobatrachidae

The Ceratobatrachidae are a family of frogs^[1][2] found in the Malay Peninsula, Borneo, the Philippines, Palau, Fiji, New Guinea, and the Admiralty, Bismarck, and Solomon Islands.^[1]

Ceratobatrachidae
Cornufer guentheri
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Amphibia
Order:	Anura
Clade:	Ranoidea
Family:	Ceratobatrachidae Boulenger, 1884
Type genus
Ceratobatrachus Boulenger, 1884
Genera
5, see text.
Synonyms
Ceratobatrachinae

Taxonomy

Ceratobatrachidae was formerly treated as a subfamily (i.e., Ceratobatrachinae) in the family Ranidae (true frogs), but have now been re-classified as a separate family. The following genera are recognised:^[1]

• Subfamily Alcalinae Brown, Siler, Richards, Diesmos, and Cannatella, 2015
  • Alcalus (5 species)
• Subfamily Ceratobatrachinae Boulenger, 1884
  • Cornufer Tschudi, 1838 (> 50 species)
  • Platymantis Günther, 1858 (> 30 species)
• Subfamily Liuraninae Fei, Ye, and Jiang, 2010
  • Liurana Dubois, 1987 (4 species)

Formerly, the following genera were also recognized in the family Ceratobatrachidae, but have now been merged into the genera above.

• Batrachylodes Boulenger, 1887 (8 species)
• Palmatorappia Ahl, 1927 (1 species)
• Ceratobatrachus Boulenger, 1884 (1 species)
• Discodeles Boulenger, 1918 (5 species)

Distribution

Ceratobatrachidae is distributed across Island Southeast Asia,^[3] as well as in the Eastern Himalayas.

Genus Liurana

• Eastern Himalayas: 4 species

Genus Alcalus

• Borneo: 2 species (Alcalus baluensis and Alcalus rajae)
• Palawan: 1 species (Alcalus mariae)

Genus Platymantis

• Philippines:

Genus Cornufer

• Palau: 1 species
• Maluku: 3–4 species
• New Guinea: 6–8 species
• Bismarck Archipelago: 18–22 species
• Solomon Islands: 20–25 species
• Fiji: 2 species

Life history

All Ceratobatrachidae lay eggs outside of water and undergo direct development where eggs hatch directly into froglets, without free-living tadpole stages.^[4]