Geomitridae

This article is about a mollusc family. For the insect family, see Geometridae.

Geomitridae is a taxonomic family of small to medium-sized air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Helicoidea.^[1][2]

Geomitridae
Cernuella virgata
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
Order:	Stylommatophora
Superfamily:	Helicoidea
Family:	Geomitridae Boettger, 1909
Type genus
Geomitra Swainson, 1840
Genera
See text

Anatomy

The family is characterized by the presence of a free right ommatophoral retractor (passing outside the peni-oviducal angle) and a double stimulatory apparatus.^[1] A free right ommatophoral retractor has been linked to adaptation to xeric habitats.^[3]

Helicella macedonica

Taxonomy

The family Geomitridae is subdivided in the following subfamilies (according to Razkin et al., 2015^[1]). Genera with the family Geomitridae include:^[1][4]

Family Geomitridae

• Not belonging to a subfamily:
  • Keraea Gude, 1911
  • Peridotitea Torres Alba, Holyoak, D. T., Holyoak, G. A., Vázquez Toro & Ripoll, 2018

• Subfamily Geomitrinae
  • tribe Cochlicellini Schileyko, 1972
    • Cochlicella Férussac, 1821
    • Monilearia Mousson, 1872
    • Obelus W. Hartmann, 1842
    • Ripkeniella Hutterer & E. Gittenberger, 1998
  • tribe Geomitrini C. Boettger, 1909 - synonym: Ochthephilinae Zilch, 1960 (n.a.)
    • Actinella R. T. Lowe, 1852^[5]
    • Callina R. T. Lowe, 1855
    • Caseolus R. T. Lowe, 1852^[5]
    • Discula R. T. Lowe, 1852^[5]
    • Disculella Pilsbry, 1895^[5]
    • Domunculifex Brozzo, De Mattia, Harl & Neiber, 2020
    • Geomitra Swainson, 1840^[5]
    • Helicomela R. T. Lowe, 1855
    • Heterostoma W. Hartmann, 1843:^[5] synonym of Steenbergia Mandahl-Barth, 1950 (invalid: junior homonym of Heterostoma de Filippi, 1837 [Platyhelminthes])
    • Hystricella R. T. Lowe, 1855^[5]
    • Lemniscia R. T. Lowe, 1855^[5]
    • Moreletina de Frias Martins, 2002^[5]
    • Plebecula R. T. Lowe, 1852
    • Pseudocampylaea L. Pfeiffer, 1877^[5]
    • Serratorotula Groh & Hemmen, 1986^[5]
    • Spirorbula R. T. Lowe, 1852^[5]
    • Steenbergia Mandahl-Barth, 1950
    • Testudodiscula Brozzo, De Mattia, Harl & Neiber, 2020
    • Wollastonaria De Mattia, Neiber & Groh, 2018
  • tribe Ponentinini Razkin et al., 2015
    • Ponentina P. Hesse, 1921

• subfamily Helicellinae
  • tribe Cernuellini Schileyko, 1991
    • Alteniella Clerx & Gittenberger, 1977
    • Cernuella Schlüter, 1838
    • Microxeromagna Ortiz de Zárate López, 1950
    • Xeroplana Monterosato, 1892
    • Xerosecta Monterosato, 1892
  • tribe Helicellini Ihering, 1909 - synonym: Jacostidae Pilsbry, 1948 (inv.)
    • Backeljaia Chueca et al., 2018
    • Candidula Kobelt, 1871
    • Helicella Férussac, 1821
    • Orexana Chueca et al., 2018
    • Xerogyra Monterosato, 1892
    • Xeroleuca Kobelt, 1877
    • Xeroplexa Monterosato, 1892
    • Xerotricha Monterosato, 1892
    • Zarateana Chueca et al., 2018
  • tribe Helicopsini H. Nordsieck, 1987
    • Helicopsis Fitzinger, 1833
    • Pseudoxerophila Westerlund, 1879
    • Xerolenta Monterosato, 1892
    • Xeromunda Monterosato, 1892
    • Xeropicta Monterosato, 1892
  • tribe Plentuisini Razkin et al., 2015
    • Plentuisa Puente & Prieto, 1992
  • tribe Trochoideini H. Nordsieck, 1987
    • Trochoidea T. Brown, 1827
    • Xerocrassa Monterosato, 1892