Granofilosea

Granofilosea is a class of cercozoan protists in the subphylum Reticulofilosa.^[2] Out of the three groups that were traditionally considered heliozoans: the heliomonads, gymnosphaerids and desmothoracids, the latter were recently grouped into this new class.

Granofilosea
Clathrulina elegans
Scientific classification
Domain:	Eukaryota
Clade:	Sar
Clade:	Rhizaria
Phylum:	Cercozoa
Subphylum:	Reticulofilosa
Class:	Granofilosea Cavalier-Smith & Bass 2009[1]
Orders
Desmothoracida Cryptofilida Limnofilida Leucodictyida

Phylogeny and taxonomy

Phylogeny

The following cladogram shows the results of a 2011 phylogenetic analysis:^[1]

	Monadofilosa Granofilosea novel clade Gran-5 Desmothoracida novel clade Gran-4 novel clades Gran-2 & 3 Cryptofilida novel clade Gran-1 Nanofilidae novel clade Gran-6 Mesofilidae Leucodictyida Limnofilida
	Chlorarachnea

Taxonomy

• Class Granofilosea Cavalier-Smith & Bass 2009^[3][4]
  • Family ?Microgromiidae De Saedeleer 1934
    • Genus Apogromia de Saedeleer 1934
    • Genus Belaria de Saedeleer 1934
    • Genus Heterogromia de Saedeleer 1934
    • Genus Microgromia Hertwig & Lesser 1874
    • Genus Paralieberkuehnia de Saedeleer 1934
  • Order ?Axomonadida [Exonucleoaxoplastiales]
    • Family Tetradimorphidae Febvre-Chevalier & Febvre 1984
      • Genus Tetradimorpha Hsiung 1927
  • Order Desmothoracida Honigberg et al. 1964 [Cryptaxohelida; Desmothoraca Hertwig & Lesser 1874; Clathrulinida Starobogatov 1980]
    • Family Clathrulinidae Claus 1874
      • Genus Actinosphaeridium Zacharias 1893
      • Genus Cienkowskya Schaudinn 1896 non Regel & Rach 1859 non Solms 1867
      • Genus Clathrulina Cienkowski 1867
      • Genus Elaeorhanis Greeff 1873
      • Genus Hedriocystis Hertwig & Lesser 1874
      • Genus Penardiophrys Mikrjukov 2000
  • Order Cryptofilida Cavalier-Smith & Bass 2009
    • Family Mesofilidae Cavalier-Smith & Bass 2009
      • Genus Mesofila Cavalier-Smith & Bass 2009
    • Family Nanofilidae Cavalier-Smith & Bass 2009
      • Genus Nanofila Bass et al. 2009
  • Order Limnofilida Cavalier-Smith & Bass 2009
    • Family Limnofilidae Cavalier-Smith & Bass 2009
      • Genus Limnofila Cavalier-Smith & Bass 2009
  • Order Leucodictyida Cavalier-Smith 1993 emend. 2003
    • Family Massisteriidae Cavalier-Smith 1993
      • Genus Massisteria Larsen & Patterson 1990
      • Genus Minimassisteria Arndt & Cavalier-Smith 2011
    • Family Leucodictyidae Cavalier-Smith 1993 [Reticulamoebidae]
      • Genus Leucodictyon Grell 1991
      • Genus Reticulamoeba Grell 1994
      • Genus Thalassomyxa Grell 1985