Graulia

Graulia is an extinct genus of lobe-finned fish from the Middle Triassic (Ladinian) of eastern France. It contains a single species, G. branchiodonta, and is represented by some of the most well preserved fossils of any Mesozoic coelacanth. Discovered in the Muschelkalk, two specimens are known, both of which are 16 cm long juveniles.^[1]

Graulia Temporal range: Ladinian PreꞒ Ꞓ O S D C P T J K Pg N ↓
Holotype (top) and referred (bottom) specimens
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Actinistia
Order:	Coelacanthiformes
Suborder:	Latimerioidei
Family:	†Mawsoniidae
Genus:	†Graulia Manuelli et al., 2024
Species:	†G. branchiodonta
Binomial name
†Graulia branchiodonta Manuelli et al., 2024

Etymology

The genus name Graulia is a reference to Graoully, a mythical dragon from the folklore of Lorraine, France, where the specimens were found. The species name branchiodonta, comes from the Greek words βράγχια "gills" and ὀδούς, ὀδόντος "tooth", the latter which refers to the large teeth of the ceratobranchials.^[1]

Description

Skeletal reconstruction

Graulia had very small teeth in its jaws, but the ceratobranchials contained much larger teeth that would have functioned similarly to the gill rakers seen in ray-finned fish. As a result, Graulia is inferred to have been a suction feeder. Enlarged sensory canals in the skull suggest strong sensitivity to minimal water changes. This, in addition to the streamlined skull and relatively large fins, support that Graulia lived a highly active predatory lifestyle.^[1]

Gallery

• 
  Micro CT scan of MHNG GEPI V5787 (holotype)
• 
  Micro CT scan of MHNG GEPI V5788 (referred)
• 
  Skull diagram