Gulella

Gulella is a genus of very small air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Streptaxidae.^[2]

Gulella
Two different views of two shells of Gulella io
Scientific classification
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
Order:	Stylommatophora
Superfamily:	Streptaxoidea
Family:	Streptaxidae
Genus:	Gulella Pfeiffer, 1856[1]
Type species
Pupa menkeana L. Pfeiffer, 1853
Synonyms
Diaphera (Huttonella) L. Pfeiffer, 1856 (superseded combination) Ennea (Gulella) L. Pfeiffer, 1856 (original rank) Ennea (Huttonella) L. Pfeiffer, 1856 (original combination) Ennea (Paucidentata) (spelling error) Ennea (Uniplicaria) L. Pfeiffer, 1856 (superseded combination) Gulella (Gulella) L. Pfeiffer, 1856· accepted, alternate representation Gulella (Huttonella) L. Pfeiffer, 1856· accepted, alternate representation Gulella (Maurennea) Schileyko, 2000· accepted, alternate representation Gulella (Molarella) Connolly, 1922· accepted, alternate representation Gulella (Paucidentina) E. von Martens, 1897· accepted, alternate representation Gulella (Plicigulella) Pilsbry, 1919· accepted, alternate representation Gulella (Tortigulella) Pilsbry, 1919· accepted, alternate representation Gulella (Uniplicaria) E. von Martens, 1895 (preoccupied name) Gulella (Uniplicaria) L. Pfeiffer, 1856· accepted, alternate representation Gulella (Wilmattina) Pilsbry & Cockerell, 1933· accepted, alternate representation Gulella (Zulugulella) Rowson & Herbert, 2016· accepted, alternate representation Huttonella L. Pfeiffer, 1856 Maurennea Schileyko, 2000 Ptychotrema (Wilmattina) Pilsbry & Cockerell, 1933 (original combination)

Until recently, species which are now in the genus Costigulella were included here.

Distribution

Distribution of the genus Gulella include:

• Afrotropical:^[3] Uganda (20 species)^[4]
• Arabia^[3]
• Madagascar^[3]
• Comoros^[3]
• the Seychelles^[3]
• Mauritius^[3]

Species

Species within the large genus Gulella include:

• Gulella aberdarensis Preston, 1913
• Gulella acutidens (O. Boettger, 1905)
• Gulella adami van Bruggen, 1994
• Gulella aequidentata (E. A. Smith, 1890)
• Gulella afrooccidentalis Verdcourt, 2004
• Gulella albersi (L. Pfeiffer, 1855)
• Gulella albinus van Bruggen & Van Goethem, 1999
• Gulella albogilva Germain, 1934
• Gulella aliciae (Melvill & Ponsonby, 1907)
• Gulella alleni Verdcourt, 1974
• Gulella alutacea Connolly, 1932
• Gulella amaniensis Verdcourt, 1953
• Gulella ambalaniranae Emberton, 2001
• Gulella ambanikelia Emberton, 2001
• Gulella ambodipelomosiae Emberton, 2002^[5]
• Gulella ambrensis Emberton, 2001
• Gulella amicta (E. A. Smith, 1890)
• Gulella analamerae Emberton, 2001
• Gulella andreana Fischer-Piette, Blanc, F. & Vukadinovic, 1974
• Gulella andriantanteliae^[6]
• Gulella ankaranaensis Fischer-Piette, Blanc, C.P., Blanc, F. & Salvat, 1994
• Gulella annibiodiversitatis van Bruggen, 2011
• Gulella antelmeana Peile, 1936
• Gulella antongilae Emberton, 2001
• Gulella appletoni van Bruggen, 1975
• Gulella aprosdoketa Connolly, 1939
• Gulella aranearum van Bruggen, 1986
• Gulella argoudi Griffiths, 2000
• Gulella arnoldi (Sturany, 1898)
• Gulella arthuri (Dautzenberg, 1890)
• Gulella atewana de Winter, 1996
• Gulella aversostriata Verdcourt, 1985
• Gulella azeitonae D. Holyoak & G. Holyoak, 2020
• Gulella baccata (Preston, 1913)
• Gulella bancoensis de Winter, 1996
• Gulella barbarae Connolly, 1929
• Gulella barnardi van Bruggen, 1965
• Gulella bassetti Burnup, 1925
• Gulella beandreana Emberton, 2001^[7]
• Gulella bebokae Emberton, 2001
• Gulella bemarahae Emberton, 2001
• Gulella bemoka Emberton, 2001
• Gulella benetecta Connolly, 1930
• Gulella benjamini Emberton & Pearce, 2000
• Gulella bernardi van Bruggen & Van Goethem, 1997
• Gulella betamponae Emberton, 2002^[5]
• Gulella bicolor (T. Hutton, 1834)
• Gulella bitzeense Connolly, 1922
• Gulella bobaombiae Emberton, 2001
• Gulella bolocoensis Ortiz de Zárate López & Ortiz de Zárate Rocandio, 1956
• Gulella bomolensis Verdcourt, 1953
• Gulella bomvana Cole & Herbert, 2009
• Gulella bondi Verdcourt, 1962
• Gulella bouchardi Fischer-Piette, Blanc, F. & Vukadinovic, 1974
• Gulella boucheti Fischer-Piette, Blanc, C.P., Blanc, F. & Salvat, 1994
• Gulella bowkerae (Melvill & Ponsonby, 1892)
• Gulella brevicula (Morelet, 1882)
• Gulella brevis (Thiele, 1911)
• Gulella bruggeni Cole & Herbert, 2009
• Gulella burnupi (Melvill & Ponsonby, 1897)
• Gulella burungaensis (Preston, 1913)
• Gulella bushmanensis Burnup, 1926
• Gulella cacosystata Germain, 1934
• Gulella cacuminata Ortiz de Zárate López & Ortiz de Zárate Rocandio, 1956
• Gulella cairnsi (Melvill & Ponsonby, 1897)
• Gulella callosa (Morelet, 1881)
• Gulella calopasa (Melvill & Ponsonby, 1903)
• Gulella camerani (Pollonera, 1906)
• Gulella cancellata Connolly, 1922
• Gulella candela Connolly, 1922
• Gulella candidula (Morelet, 1889)
• Gulella capmini Emberton, 2001
• Gulella cara Pilsbry, 1919^[8]
• Gulella carea (Preston, 1913)
• Gulella caroli (Kobelt, 1913)
• Gulella carpenteri Connolly, 1931
• Gulella caryatis (Melvill & Ponsonby, 1898)
• Gulella cavidens (E. von Martens, 1876)
• Gulella ceciliae van Bruggen, 1971
• Gulella celestinae Emberton, 2001
• Gulella cerea (Dunker, 1848)
• Gulella chapini Pilsbry, 1919
• Gulella cheranganiensis Germain, 1934
• Gulella chi Burnup, 1926
• Gulella claustralis Connolly, 1939
• Gulella claustrum (Preston, 1913)
• Gulella coarctata (D'Ailly, 1910)
• Gulella coarti (Dautzenberg & Germain, 1914)
• Gulella coeni (Preston, 1913)
• Gulella collicola van Bruggen, 1966
• Gulella columella (E. A. Smith, 1903)
• Gulella columna Emberton, 2001
• Gulella columnella (Melvill & Ponsonby, 1901)
• Gulella commoda (E. A. Smith, 1903)
• Gulella comorensis (E. von Martens, 1876)
• Gulella conicodentata K. L. Pfeiffer, 1952
• Gulella connollyi (Melvill & Ponsonby, 1909)
• Gulella conradti (E. von Martens, 1895)
• Gulella consanguinea (E. A. Smith, 1890)
• Gulella consobrina (Ancey, 1892)
• Gulella consociata (E. A. Smith, 1890)
• Gulella constricta Emberton, 2001
• Gulella contingens Burnup, 1925
• Gulella contraria Connolly, 1932
• Gulella copiosa (Preston, 1913)
• Gulella corneola (Morelet, 1886)
• Gulella costellata (Morelet, 1882)
• Gulella crassidens (L. Pfeiffer, 1859)
• Gulella crassilabris (Craven, 1881)
• Gulella craterodon (Melvill & Ponsonby, 1903)
• Gulella cruciata (E. von Martens, 1900)
• Gulella crux van Bruggen, 2011
• Gulella crystallum (Morelet, 1848)
• Gulella cupula van Bruggen & Van Goethem, 1999
• Gulella curvicolumella (Preston, 1913)
• Gulella curvilamella (E. A. Smith, 1890)
• Gulella cyclochilus Degner, 1934
• Gulella cylindrica (L. Pfeiffer, 1852)
• Gulella daedalea (Melvill & Ponsonby, 1903)
• Gulella darglensis (Melvill & Ponsonby, 1908)
• Gulella dartevillei Verdcourt, 1962
• Gulella dautzenbergi Connolly, 1928
• Gulella davisae Herbert, 2016
• Gulella decaryi Fischer-Piette & Vukadinovic, 1974
• Gulella decussatula (Preston, 1913)^[4]
• Gulella dejae Bursey & Herbert, 2004
• Gulella delicatula (L. Pfeiffer, 1857)
• Gulella dentiens (Morelet, 1883)
• Gulella devia Connolly, 1931
• Gulella deviae Herbert, 2006^[9]
• Gulella dextrorsa Burnup, 1925
• Gulella diabensis Connolly, 1939
• Gulella diodon (Morelet, 1882)
• Gulella discrepans (Sturany, 1898)
• Gulella disseminata (Preston, 1913)
• Gulella distincta (Melvill & Ponsonby, 1893)
• Gulella dohrni (E. A. Smith, 1882)
• Gulella dolichos Verdcourt, 1985
• Gulella doliolum (Morelet, 1867)
• Gulella dorri (Dautzenberg, 1890)
• Gulella drakensbergensis (Melvill & Ponsonby, 1893)
• Gulella duncani Connolly, 1930
• Gulella dunkeri (L. Pfeiffer, 1856)
• Gulella dupuisi Connolly, 1922
• Gulella dupuyana (Crosse, 1876)
• Gulella ectodentata van Bruggen & Van Goethem, 1999
• Gulella elliptica (Melvill & Ponsonby, 1898)
• Gulella enneodon Connolly, 1922
• Gulella eoryi Verdcourt, 2004
• Gulella erugo Rowson, Foster, Seddon & Tattersfield, 2019
• Gulella escalarai Ortiz de Zárate López & Ortiz de Zárate Rocandio, 1956
• Gulella eussoensis (Preston, 1913)
• Gulella euthymia (Melvill & Ponsonby, 1893)
• Gulella excavata (E. von Martens, 1892)
• Gulella excruciata Connolly, 1931
• Gulella exogonia (E. von Martens, 1895)
• Gulella expatriata (Preston, 1910)
• Gulella falconi Burnup, 1925
• Gulella farquhari (Melvill & Ponsonby, 1895)
• Gulella fernandensis Ortiz de Zárate López & Ortiz de Zárate Rocandio, 1956
• Gulella filix Connolly, 1922
• Gulella fischerpiettei Emberton, 2001
• Gulella formosa (Melvill & Ponsonby, 1898)
• Gulella fortidentata (E.A. Smith, 1890)
• Gulella fotobohitrae Emberton, 2001
• Gulella foveolata Preston, 1913
• Gulella framesi Burnup, 1926
• Gulella fraudator Connolly, 1939^[9]
• Gulella funerea (Preston, 1913)
• Gulella galactochila (Crosse, 1885)
• Gulella gallorum Fischer-Piette, Blanc, F. & Salvat, 1975
• Gulella garambae van Bruggen & Van Goethem, 1999
• Gulella genialis (Melvill & Ponsonby, 1903)
• Gulella gestroi (Germain, 1915)
• Gulella girardi (Kobelt, 1904)
• Gulella glabra (Morelet, 1882)
• Gulella godfreyi Burnup, 1925
• Gulella gouldi (L. Pfeiffer, 1856)
• Gulella greenwayi Verdcourt, 1953
• Gulella griffithsi Emberton, 2001
• Gulella guilielmi van Bruggen & Van Goethem, 1998
• Gulella gwendolinae (Preston, 1910)
• Gulella hadroglossa Herbert, 2016
• Gulella hafa Emberton, 2001
• Gulella hafahafa Emberton, 2001
• Gulella hamerae Bursey & Herbert, 2004
• Gulella handeiensis Verdcourt, 1953
• Gulella harriesi Burnup, 1926
• Gulella haullevillei (Dautzenberg & Germain, 1914)
• Gulella hector (Preston, 1913)
• Gulella helichrysophila Germain, 1934
• Gulella hemmingi Verdcourt, 1963
• Gulella herberti van Bruggen, 2004
• Gulella heteromphala Pilsbry, 1919
• Gulella hildae van Bruggen, 2001
• Gulella himerothales (Melvill & Ponsonby, 1903)
• Gulella hoqensis Neubert, 2004
• Gulella hordeum (Morelet, 1879)
• Gulella hughscotti Verdcourt, 1980
• Gulella humbloti (Morelet, 1886)
• Gulella impedita Connolly, 1922
• Gulella inconspicua (Thiele, 1911)
• Gulella incurvidens van Bruggen, 1972
• Gulella infans (Craven, 1881)
• Gulella infrendens (E. von Martens, 1866)
• Gulella ingloria (Preston, 1913)
• Gulella inhluzaniensis (Burnup, 1914)
• Gulella inobstructa van Bruggen, 1965
• Gulella insolita (E. A. Smith, 1903)
• Gulella instabilis (Sturany, 1898)
• Gulella insularis (Girard, 1894)
• Gulella insulincola van Bruggen, 1975
• Gulella intradentata (Preston, 1913)
• Gulella intrusa Verdcourt, 1956
• Gulella io Verdcourt, 1974
• Gulella iridescens (Preston, 1913)
• Gulella isipingoensis (Sturany, 1898)
• Gulella isseli (Paladilhe, 1872)
• Gulella jaominai Emberton, 2001
• Gulella jod (Preston, 1910)
• Gulella johannae van Bruggen, 2006
• Gulella johannesburgensis (Melvill & Ponsonby, 1907)
• Gulella jongkindi de Winter, 1996
• Gulella josephinae Emberton, 2001
• Gulella juxtidens (Melvill & Ponsonby, 1899)
• Gulella kelibea Emberton, 2001
• Gulella kelimolotra Emberton, 2002^[5]
• Gulella kendrae Emberton & Griffiths, 2009^[7]
• Gulella kidundae Adam, 1965
• Gulella kigeziensis (Preston, 1913)
• Gulella kohllarseni (Haas, 1936)
• Gulella kosiensis (Melvill & Ponsonby, 1908)
• Gulella kraussi (L. Pfeiffer, 1856)
• Gulella kulalensis Verdcourt, 1962
• Gulella labiotuberculata Connolly, 1942
• Gulella lacuna (Preston, 1911)
• Gulella laevigata (Dohrn, 1865)
• Gulella laevorsa Burnup, 1925
• Gulella lambda Degner, 1934
• Gulella lamyi (Dautzenberg & Germain, 1914)
• Gulella landianiensis (Dautzenberg, 1908)
• Gulella langei Rowson, Foster, Seddon & Tattersfield, 2019
• Gulella laninifia Emberton, 2002^[5]
• Gulella laqueus (Preston, 1913)
• Gulella larva (Morelet, 1877)
• Gulella latimerae Bursey & Herbert, 2004
• Gulella lawrencei van Bruggen, 1964
• † Gulella leakeyi Verdcourt, 1963
• Gulella lendix (E. A. Smith, 1890)
• Gulella lessensis Pilsbry, 1919
• Gulella leucocion Connolly, 1929
• Gulella lievrouwi van Bruggen & Van Goethem, 1999
• Gulella lima (Preston, 1913)
• Gulella lincolni Emberton & Griffiths, 2009^[7]
• Gulella lindae Herbert, 2006^[9]
• Gulella linguidens Connolly, 1939
• Gulella lissophanes (Melvill & Ponsonby, 1892)
• Gulella lohabea Emberton, 2001
• Gulella lornae Verdcourt, 1952
• Gulella loveridgei van Bruggen, 1996
• Gulella lubeti Fischer-Piette, Blanc, C.P., Blanc, F. & Salvat, 1994
• Gulella lubrica (Morelet, 1881)
• Gulella ludwigi Verdcourt & Venmans, 1953
• Gulella magnifica Emberton, 2001
• Gulella magnolia (Connolly, 1912)
• Gulella magnorchida Emberton, 2001
• Gulella mahafinaratra Emberton, 2001
• Gulella mahagaga Emberton, 2001
• Gulella mahia Emberton, 2001
• Gulella malasangiensis (Preston, 1913)
• Gulella mamellensis Griffiths, 2000
• Gulella manomboae Emberton, 2001
• Gulella mariae (Melvill & Ponsonby, 1892)
• Gulella marionae (Preston, 1910)
• Gulella maritzburgensis (Melvill & Ponsonby, 1893)
• Gulella marojejyae Emberton, 2001
• Gulella masoalae Emberton, 2001
• Gulella masis iensis Pilsbry, 1919
• Gulella matavymolotra Emberton, 2002^[5]
• Gulella matumbiensis Verdcourt, 1993
• Gulella mayottensis Connolly, 1925
• † Gulella mbili Pickford, 2009
• Gulella megapex Neubert, 2004
• Gulella melvilli (Burnup, 1914)
• Gulella meneleki (Preston, 1910)
• Gulella menkeana (L. Pfeiffer, 1853)
• Gulella menkhorsti van Bruggen, 2011
• Gulella meruensis (D'Ailly, 1910)
• Gulella mfongosiensis Burnup, 1925
• Gulella miaranoniae Emberton, 2001
• Gulella miaryi Fischer-Piette & Bedoucha, 1964
• Gulella micans K. L. Pfeiffer, 1952
• Gulella michellae Emberton, 2001
• Gulella microdina (Morelet, 1883)
• Gulella microdon (Morelet, 1860)
• Gulella microrutshuruensis van Bruggen, 1995
• Gulella microstriata Emberton, 2001
• Gulella mihomehia Emberton, 2001
• Gulella mikenoensis (Preston, 1913)
• Gulella miniata (F. Krauss, 1848)
• Gulella minuscula Emberton & Pearce, 2000^[5]
• Gulella minuta (Morelet, 1889)
• Gulella minutissima (Thiele, 1911)
• Gulella mitsikia Emberton, 2001
• Gulella mkuu Rowson, Seddon & Tattersfield, 2009
• Gulella modioliformis (Morelet, 1877)
• † Gulella moja Pickford, 2009
• Gulella mongolae Ortiz de Zárate Lopez & Ortiz de Zárate Rocandio, 1956
• Gulella monodon (Morelet, 1873)
• Gulella montium (D'Ailly, 1910)
• Gulella mooiensis (Burnup, 1914)
• Gulella msoalae Emberton, 2001
• Gulella multidentata (Sturany, 1898)
• Gulella munita (Melvill & Ponsonby, 1892)
• Gulella nakamaroa Emberton, 2001
• Gulella namorokae Emberton, 2001
• Gulella natalensis (Craven, 1881)
• Gulella ndibo Cole & Herbert, 2009
• Gulella nemoralis (Germain, 1915)
• Gulella nepia Connolly, 1925
• Gulella newmani Bursey & Herbert, 2004
• Gulella newtoni (E. A. Smith, 1890)
• Gulella ngorogoroensis Verdcourt, 1985
• Gulella nifikelia Emberton, 2001
• † Gulella nne Pickford, 2009
• Gulella noltei (Boettger, 1898)
• Gulella nosybei Emberton, 2001
• Gulella nuchalis van Bruggen, 2011
• Gulella nyikaensis (Preston, 1913)
• Gulella nyiroensis (Preston, 1913)
• Gulella obani Oke, 2007^[10]
• Gulella obovata (L. Pfeiffer, 1855)
• Gulella obstructa van Bruggen, 1965
• Gulella odhneriana Dupuis, 1923
• Gulella odietei Oke, 2013
• Gulella ogbeifuni Oke, 2013
• Gulella olkokolae W. Adam, 1965
• Gulella opoboensis (Preston, 1914)
• Gulella orchida Emberton, 2001
• Gulella orientalis Connolly, 1929
• Gulella oryza (Morelet, 1882)
• Gulella ovalis (Thiele, 1911)
• Gulella ovularis (Morelet, 1886)
• Gulella pangumana Connolly, 1928
• Gulella papyracea (Preston, 1913)
• Gulella paucidens Verdcourt, 1953
• Gulella pearcei Emberton, 2001
• Gulella peculiaris (E. A. Smith, 1890)
• Gulella penningtoni Burnup, 1925
• Gulella pentheri (Sturany, 1898)
• Gulella pentodon (Morelet, 1889)
• Gulella percivali (Preston, 1913)
• Gulella pergrata (Preston, 1913)
• Gulella perissodonta (Sturany, 1898)
• Gulella perlata Connolly, 1922
• Gulella perplexa Connolly, 1939
• Gulella perspicua (Melvill & Ponsonby, 1893)
• Gulella perspicuaeformis (Sturany, 1898)
• Gulella pervitrea (Preston, 1913)
• Gulella petitboucheti Emberton, 2001
• Gulella pfeifferi (Krauss in Kuster, 1844)
• Gulella phanerodon (Morelet, 1888)
• Gulella phragma (Melvill & Ponsonby, 1907)
• Gulella phyllisae Burnup, 1925
• Gulella planidens (E. von Martens, 1892)
• Gulella planti (L. Pfeiffer, 1856) – Plant's gulella snail
• Gulella platycostata Neubert, 2004
• Gulella plicigera (Morelet, 1886)
• Gulella polita (Melvill & Ponsonby, 1893)
• Gulella polloneriana Pilsbry, 1919
• Gulella pondoensis Connolly, 1939
• Gulella ponsonbyi (Burnup, 1914)
• Gulella porcina Connolly, 1930
• Gulella poutrini (Germain, 1918)
• Gulella praelonga Connolly, 1922
• Gulella premnodes (Sturany, 1902)
• Gulella prestoni Connolly, 1922
• Gulella pretoriana Connolly, 1932
• Gulella princei (Preston, 1911)
• Gulella protruda Neubert & Frank, 1996
• Gulella proxima van Bruggen, 2014
• Gulella pseudandreana Emberton, 2001
• Gulella pseudolkokolae Verdcourt, 2004
• Gulella puella Connolly, 1929
• Gulella pulchella (Melvill & Ponsonby, 1893)
• Gulella pusilla (Morelet, 1881)
• Gulella puzeyi Connolly, 1939
• Gulella queketti (Melvill & Ponsonby, 1896)
• Gulella raffrayi (Bourguignat, 1883)
• Gulella rakotoarisoni Emberton, 2001
• Gulella ranomasina Emberton, 2001
• Gulella razafyi Emberton, 2001
• Gulella rectangularis (Preston, 1913)
• Gulella reeae Emberton & Pearce, 2000
• Gulella reesi (Preston, 1914)
• Gulella reniformis (Preston, 1913)
• Gulella rewgidensis Neubert, 2004
• Gulella rhodesiana (Connolly, 1912)
• Gulella roberti (Preston, 1910)
• Gulella rogersi (Melvill & Ponsonby, 1898)
• Gulella rubinsterni Fischer-Piette, Blanc, C.P., Blanc, F. & Salvat, 1994
• Gulella rugosa Emberton, 2001
• Gulella rumpiana Connolly, 1932
• Gulella runa Rowson, Foster, Seddon & Tattersfield, 2019
• Gulella ruthae Emberton, 2002^[5]
• Gulella rutshuruensis Pilsbry, 1919
• Gulella ruwenzoriensis van Bruggen & van Goethem, 1999^[4]
• Gulella sahia Emberton, 2002^[5]
• Gulella salpinx Herbert, 2002^[11]
• Gulella sandersoni Connolly, 1942
• Gulella sankuruensis Dartevelle-Puissant, 1936
• Gulella satisfacta Fischer-Piette, Blanc, C.P., Blanc, F. & Salvat, 1994
• Gulella schweinfurthi (Thiele, 1910)
• Gulella sellae (Pollonera, 1906)
• Gulella seneciophila Germain, 1934
• Gulella separata (Sturany, 1898)
• Gulella sesamum (Morelet, 1883)
• Gulella sexdentata (E. von Martens, 1869)
• Gulella shabae Adam & Van Goethem, 1978
• Gulella shoaensis Verdcourt, 1985
• Gulella sibasana Connolly, 1922
• Gulella simplicima (Preston, 1911)
• † Gulella sita Pickford, 2009
• Gulella socialis Pilsbry, 1919
• Gulella socotrensis Neubert, 2004
• Gulella soror (E. A. Smith, 1890)
• Gulella soulaiana Fischer-Piette, 1973^[7]
• Gulella spreta (Morelet, 1883)
• Gulella stolidodea Degner, 1934
• Gulella streptostelopsis Bruggen, 2007^[12]
• Gulella strictilabris (Ancey, 1898)
• Gulella stylodon (E. von Martens, 1876)
• Gulella suavissima (Preston, 1913)
• Gulella subflavescens (E. A. Smith, 1890)
• Gulella subframesi Connolly, 1929
• Gulella subhyalina (E. A. Smith, 1890)
• Gulella subkraussi Connolly, 1932
• Gulella subringens (Crosse, 1886)
• Gulella subsellae Haas, 1936
• Gulella sursum van Bruggen, 2001
• Gulella swaziensis Connolly, 1932
• Gulella sylvia (Melvill & Ponsonby, 1903)
• Gulella syngenes (Preston, 1913)
• Gulella systemanaturae van Bruggen, 2008^[13]
• Gulella taitensis Verdcourt, 1963
• † Gulella tano Pickford, 2009
• Gulella taolantehezana Emberton, 2002^[5]
• † Gulella tatu Pickford, 2009
• Gulella tendronia Emberton, 2001
• Gulella tharfieldensis (Melvill & Ponsonby, 1893)
• Gulella thompsoni K. C. Emberton, Slapcinsky, Campbell, Rakotondrazafy, Andriamiarison & J. D. Emberton, 2010^[6]
• Gulella tietzae Cole & Herbert, 2009
• Gulella titania Connolly, 1928
• Gulella tomlini (Preston, 1911)
• Gulella tracheia Rowson, 2007
• Gulella translucida K. L. Pfeiffer, 1952
• Gulella transnominata Connolly, 1925
• Gulella triglochis (Melvill & Ponsonby, 1903)
• Gulella trigona (Morelet, 1881)
• Gulella triplicina (E. von Martens, 1895)
• Gulella tripodium Connolly, 1939
• Gulella tristaoensis Connolly, 1922
• Gulella tsara Emberton, 2001
• Gulella tsaratananae Emberton, 2001
• Gulella tudes (E. von Martens, 1895)
• Gulella udzungwensis van Bruggen, 2003
• Gulella ugandensis (E. A. Smith, 1901)
• Gulella uluguruensis Verdcourt, 1962
• Gulella umzimvubuensis Burnup, 1925
• Gulella unidentata K. L. Pfeiffer, 1952
• Gulella vakinifia Emberton, 2001
• Gulella vallaris (Melvill & Ponsonby, 1907)
• Gulella varians (E. A. Smith, 1899)
• Gulella vatosoa Emberton, 2001
• Gulella vavakelia Emberton, 2001
• Gulella ventricosa Neubert, 2004
• Gulella verdcourti van Bruggen, 1966
• Gulella vermis (Morelet, 1881)
• Gulella viae Burnup, 1925
• Gulella viatoris (Preston, 1913)
• Gulella vicina (E.A. Smith, 1899)
• Gulella virungae van Bruggen & van Goethem, 1999^[4]
• Gulella vohimarae Emberton, 2001
• Gulella vriesiana (C. M. F. Ancey, 1885)
• Gulella wahlbergi (F. Krauss, 1848)
• Gulella warrenii (Melvill & Ponsonby, 1903)
• Gulella wendalinae van Bruggen, 1975
• Gulella wilmattae (Pilsbry & Cockerell, 1933)
• Gulella woodhousei (Preston, 1913)
• Gulella xysila (Melvill & Ponsonby, 1907)
• Gulella zanaharyi Emberton, 2001
• Gulella zelota (Melvill & Ponsonby, 1907)
• Gulella zemenensis Verdcourt, 1990
• Gulella zuluensis Connolly, 1932
• Gulella sp. 1 sensu Emberton & Griffiths (2009)^[7]
• Gulella sp. 2 sensu Emberton & Griffiths (2009)^[7]
• Gulella sp. 3 sensu Emberton & Griffiths (2009)^[7]

Synonyms:

• Gulella amboniensis : synonym of Juventigulella amboniensis (Tattersfield, 1998) (original combination)
• Gulella avakubiensis Pilsbry, 1919:^[4] synonym of Avakubia avakubiensis (Pilsbry, 1919) (superseded combination)
• Gulella caryatis diabensis Connolly, 1939:^[3] synonym of Gulella diabensis Connolly, 1939 (original combination)
• Gulella conospira (Martens, 1892):^[4] synonym of Conogulella conospira (E. von Martens, 1892) (superseded combination)
• Gulella cuspidata Verdcourt, 1962: synonym of Dadagulella cuspidata (Verdcourt, 1962) (original combination)
• Gulella foliifera (E. von Martens, 1895): synonym of Primigulella foliifera (E. von Martens, 1895) (superseded combination)
• Gulella grossa (E. von Martens, 1892): synonym of Primigulella grossa (E. von Martens, 1892) (superseded combination)
• Gulella kuiperi de Winter, 2007:^[14] synonym of Silvigulella kuiperi (De Winter, 2006) (basionym)
• Gulella linguifera (Martens, 1895):^[4] synonym of Primigulella linguifera (E. von Martens, 1895) (superseded combination)
• Gulella ndamanyiluensis]] Venmans, 1956: synonym of Primigulella ndamanyiluensis (Venmans, 1956) (original combination)
• Gulella osborni Pilsbry, 1919:^[4] synonym of Silvigulella osborni (Pilsbry, 1919) (original combination)
• Gulella pupa (Thiele, 1911):^[4] synonym of Pupigulella pupa (Thiele, 1911) (superseded combination)
• Gulella selene van Bruggen & van Goethem, 1999:^[4] synonym of Dadagulella selene (van Bruggen & Van Goethem, 1999) (original combination)
• Gulella thomasseti is a synonym for Glabrennea thomasseti (Sykes, 1909)^[15]
• Gulella usambarica K. L. Pfeiffer, 1952: synonym of Gulella ludwigi Verdcourt & Venmans, 1953 (junior secondary homonym of Gulella usambarica (Craven, 1880); Gulella ludwigi Verdcourt & Venmans, 1953 is a replacement name)
• Gulella usambarica (Craven, 1880): synonym of Primigulella usambarica (Craven, 1880) (superseded combination)