Gymnosphaera

Gymnosphaera is a genus of tree ferns in family Cyatheaceae.

Gymnosphaera
Scientific classification
Kingdom:	Plantae
Clade:	Tracheophytes
Division:	Polypodiophyta
Class:	Polypodiopsida
Order:	Cyatheales
Family:	Cyatheaceae
Genus:	Gymnosphaera Blume
Type species
Gymnosphaera glabra Blume
Species
See text
Synonyms
Alsophila section Gymnosphaera (Blume) Moore Alsophila (Gymnosphaera) (Blume) Q.Xia Cyathea (Gymnosphaera) (Blume) Tindale Cyathea section Gymnosphaera (Blume) Holttum

Taxonomy

The genus was originally described by Carl Ludwig Blume in 1828.^[1] It was frequently treated by later authors as a synonym of Cyathea or Alsophila.

In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), Gymnosphaera was left as a synonym of Alsophila, as evidence to support its reliable separation was lacking.^[2] More recently, Shi-Yong Dong and collaborators proposed the revival and recircumscription of the genus based on additional phylogenetic evidence,^[3] an act subsequently accepted by other tree fern workers.^[4]

Phylogeny

As of January 2023^[update], the following species are accepted in the genus as currently circumscribed by the Checklist of Ferns and Lycophytes of the World:^[5]

Phylogeny of Gymnosphaera[6][7]

Other species include:

Gymnosphaera G. capensis (L.fil.) S.Y.Dong G. schliebenii (Reimers) S.Y.Dong Alsophila boiviniiformis (Rakotondr. & Janssen) Christenhusz 2018 G. impolita (Rakotondr. & Janssen) S.Y.Dong G. andohahelensis (Tardieu) Tardieu G. poolii (C.Chr.) S.Y.Dong G. rouhaniana (Rakotondr. & Janssen) S.Y.Dong G. salvinii (Hook.) S.Y.Dong G. hornei (Baker) Copel. Alsophila klossii (Ridl.) Tryon G. acrostichoides (Alderw.) S.Y.Dong G. schlechteri (Brause) Copel. G. olivacea (Brause) S.Y.Dong G. phlebodes (Lehnert & Coritico) S.Y.Dong G. baileyana (Domin) S.Y.Dong G. rebeccae (F.Muell.) S.Y.Dong G. annae (Alderw.) S.Y.Dong G. dimorpha (Christ) S.Y.Dong G. biformis (Rosenst.) Copel. G. scandens (Brause) S.Y.Dong G. ramispina (Hook.) Copel. G. ogurae (Hayata) Tagawa G. subdubia (Alderw.) S.Y.Dong G. khasyana (T.Moore ex Kuhn) Ching G. lurida (Blume) S.Y.Dong G. denticulata (Baker) Copel. G. bachmaensis S.Y.Dong G. podophylla (Hook.) Copel. G. salletii (Tardieu & C.Chr. ex C.Chr. & Tardieu) S.Y.Dong G. austroyunnanensis (S.G.Lu) S.G.Lu & Chun X.Li G. saxicola S.Y.Dong & Z.Y.Zuo G. gigantea (Wall. ex Hook.) J.Sm. G. metteniana (Hance) Tagawa G. andersonii (Scott ex Beddome) Ching & Wu G. glabra Blume

Gymnosphaera alticola Tardieu Gymnosphaera atropurpurea (Copel.) Copel. Gymnosphaera bonii (Christ) S.Y.Dong Gymnosphaera commutata (Mett.) S.Y.Dong Gymnosphaera henryi (Baker) S.R.Ghosh Gymnosphaera mildbraedii (Brause) S.Y.Dong Gymnosphaera nicklesii Tardieu & F.Ballard ex Tardieu Gymnosphaera ramispinoides (M.Kato) S.Y.Dong Gymnosphaera rubella (Holttum) S.Y.Dong