Hadrianus (turtle)

Hadrianus is an extinct genus of tortoise belonging to the Testudinidae^[1] found in the United States, the Yolomécatl Formation of Mexico, the Alai Beds of Kyrgyzstan and Spain^{[citation needed]} and believed to be the oldest true tortoise known.^[3] The genus is thought to be closely related to the genus Manouria.^[3] The genus may have evolved in the subtropics of Asia and subsequently migrated to North America and Europe.^[3] Evangelos Vlachos (2018) reassessed the North American species attributed to the genus, and determined only two as accepted namely H. corsoni & H. majusculus. The remaining species were identified as either junior synonyms, moved to other genera or considered nomen dubium do to incomplete fossils.^[4]

Hadrianus Temporal range: 55.4–33.9 Ma PreꞒ Ꞓ O S D C P T J K Pg N Eocene[1]
Hadrianus majusculus
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Order:	Testudines
Suborder:	Cryptodira
Clade:	Pantestudinoidea
Superfamily:	Testudinoidea
(unranked):	†Pantestudinidae
Genus:	†Hadrianus Cope, 1872[2][1]
Species
H. corsoni H. majusculus

Taxonomy

• Hadrianus corsoni (Leidy, 1871)
  • Synonyms Emys carteri Leidy, 1871; Testudo hadriana Cope, 1871; Hadrianus quadratus Cope, 1871; Hadrianus octonaria Cope, 1871; Hadrianus tumidus Hay 1908; Hadrianus robustus Gilmore, 1915; Hadrianus utahensis Gilmore, 1915; Geochelone gilmorei Auffenberg, 1974
• Hadrianus majusculus Hay, 1904

Nomen dubium

• "Hadrianus allabiatus" Cope, 1871

Moved species

• "Hadrianus schucherti" Hay, 1899 moved to Cymatholcus schucherti (Hay, 1899)