Harmonia (beetle)

Harmonia is a genus of lady beetles belonging to the subfamily Coccinellinae.

Harmonia
Asian lady beetle, Harmonia axyridis
Harmonia testudinaria
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Coleoptera
Suborder:	Polyphaga
Infraorder:	Cucujiformia
Family:	Coccinellidae
Tribe:	Coccinellini
Genus:	Harmonia Mulsant, 1846
Synonyms
Ballia Mulsant, 1853 Callineda Crotch, 1871 Leis Mulsant, 1850 Ptychanatis Crotch, 1874 Rhopaloneda Timberlake, 1943 Stictoleis Crotch, 1874

Description

The distinction of the individual species within the genus is often very difficult, and in some cases requires examination of the reproductive system. The coloring and drawing in the genus Harmonia are very variable, not only between species, but also within each species. The basic color of the elytra is usually red to orange with black spots. The patches can be linked together to form black bars. The species can reach a length of 3 to 7 mm. The rear edge of the pronotum is slightly curved. The antennae are a little shorter than the head and are composed of eleven segments. The three last segments are thickened at the tip and form a club.

List of species

• Harmonia andamanensis Poorani, 2023
• Harmonia antipodum (White, 1848) – antipodean ladybird, endemic to New Zealand
• Harmonia areolata Bielawski & Chûjô, 1968
• Harmonia axyridis (Pallas, 1771) – Asian lady beetle, originally from Nepal, China, Taiwan, Japan, introduced to Western Europe and America
• Harmonia basinotata Bielawski, 1964 – New Guinea
• Harmonia bicolor (Blackburn, 1892) – Australia
• Harmonia boothi Ślipiński, Li & Pang, 2020
• Harmonia boulardi (Mulsant, 1850)
• Harmonia conformis (Boisduval, 1835) – common spotted lady beetle, Australia, introduced in New Zealand
• Harmonia coryphaea (Guérin-Méneville, 1844)
• Harmonia crucigera (Gyllenhaal, 1808)
• Harmonia decussata (Crotch, 1874)
• Harmonia deyrollii (Crotch, 1874)
• Harmonia dimidiata (Fabricius, 1781) – India, Pakistan, Nepal, Bhutan, China, Taiwan, Japan; introduced in North America
• Harmonia eucharis (Mulsant, 1850) – India, Pakistan, Myanmar, Nepal, Himalayan Region, South China
• Harmonia expallida (Weise, 1907) – India
• Harmonia fijiensis (Sicard, 1929)
• Harmonia gilvella Fürsch, 2001
• Harmonia instabilis (Mulsant, 1850)
• Harmonia lapeyrousei (Boisduval, 1835)
• Harmonia manillana (Mulsant, 1866)
• Harmonia micra Ślipiński, Li & Pang, 2020
• Harmonia nagaii Miyatake, 1980
• Harmonia nigromarginata Bielawski & Chûjô, 1968
• Harmonia novaehebridensis (Korschefsky, 1943)
• Harmonia ocellata (Mader, 1955)
• Harmonia octomaculata (Fabricius, 1781) – India, Pakistan, Nepal, Bangladesh, Sri Lanka, Micronesia, Australia
• Harmonia oculata (Hoang, 1983)
• Harmonia ohlmusi Ślipiński, Li & Pang, 2020
• Harmonia pardalina (Gerstäcker, 1871)
• Harmonia problematica (Fürsch, 1963)
• Harmonia quadripunctata (Pontoppidan, 1763) – cream-streaked lady beetle, Europe
• Harmonia rugulosa Sicard, 1912
• Harmonia sedecimnotata (Fabricius, 1801) – India, Nepal, Southeast Asia
• Harmonia shoichii Sasaji, 1988
• Harmonia testudinaria (Mulsant, 1850) – Indonesia, New Guinea, North Australia
• Harmonia thonningii (Mulsant, 1850)
• Harmonia tobleri Ślipiński, Li & Pang, 2020
• Harmonia tricolor (Korschefsky, 1944)
• Harmonia vigintiduomaculata (Fabricius, 1792)
• Harmonia yedoensis (Takizawa, 1917) – Japan

References

• Data related to Harmonia at Wikispecies
• Media related to Harmonia at Wikimedia Commons
• Biolib
• Catalogue of Life Archived 2020-07-06 at the Wayback Machine
• Fauna Europaea