Holepyris

Holepyris is a genus of hymenopteran parasitoids in the family Bethylidae. These species use small beetle larvae as hosts for their larvae. Prior to a 2022 study that found this genus and other genera within the subfamily Epyrinae to be polyphyletic, there were 148 species in the genus Holepyris.^[1] Following a reclassification aiming at creating monophyletic genera of Epyrinae, 131 Holepyris species were placed in the genus Rysepyris (including Holepyris sylvanidis, now Rysepyris sylvanidis, a species studied as a biocontrol agent of some storage pests), and three Formosiepyris species were placed in the genus Holepyris.^[2] As a result, there are now 23 species in the genus Holepyris:

Holepyris
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Hymenoptera
Family:	Bethylidae
Genus:	Holepyris Kieffer, 1905

Species

• Holepyris africanus Kieffer, 19041
• Holepyris agraensis (Kurian, 1955)
• Holepyris algoaensis Kieffer, 1911
• Holepyris angusticollis Kieffer, 1906
• Holepyris bergevini Duchaussoy, 1916[1914]
• Holepyris bidentatus Kieffer, 1904
• Holepyris crenulatus Kieffer, 1906
• Holepyris fasciipennis Kieffer, 1906
• Holepyris fuscipennis Kieffer, 1906
• Holepyris levicollis Kieffer, 1906
• Holepyris lineatus Kieffer, 1906
• Holepyris maculipennis (Marshall, 1874)
• Holepyris marishi (Terayama, 2004)
• Holepyris natalensis Kieffer, 1913
• Holepyris orientalis Kieffer, 1906
• Holepyris punctaticollis Kieffer, 1906
• Holepyris rufitarsis Kieffer, 1906
• Holepyris rugulosus (Xu & He, 2005)
• Holepyris shiva (Terayama, 2004)
• Holepyris takasago (Terayama, 2004)
• Holepyris tibialis Kieffer, 1906
• Holepyris vietnamensis (Tsujii et al., 2015)
• Holepyris vittatus Turner, 1928