Lirainosaurinae

Lirainosaurinae is a subfamily of lithostrotian titanosaur sauropods from the Late Cretaceous of France, Spain, and Romania.

Lirainosaurines Temporal range: Late Campanian-Early Maastrichtian, 74–66 Ma PreꞒ Ꞓ O S D C P T J K Pg N
Skeletal mount of Ampelosaurus atacis
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Clade:	Dinosauria
Clade:	Saurischia
Clade:	†Sauropodomorpha
Clade:	†Sauropoda
Clade:	†Macronaria
Clade:	†Titanosauria
Clade:	†Lithostrotia
Subfamily:	†Lirainosaurinae Díez Díaz et al. 2018[1]
Genera[2][3]
†Ampelosaurus †Atsinganosaurus †Garrigatitan †Lirainosaurus †Lohuecotitan †Paludititan?[4]

Systematics

Lirainosaurinae was defined by Díez Díaz et al. (2018) as comprising "Lirainosaurus, Ampelosaurus, their common ancestor, and all of its descendants", and includes the two definitional taxa as well as Atsinganosaurus, Lohuecotitan, and Paludititan. The five genera are known from Europe and all within the timespan of the late Campanian (Lirainosaurus) to early Maastrichtian (Ampelosaurus and Atsinganosaurus). Lirainosaurinae was recovered by Diez et al. (2018) as phylogenetically intermediate between the clades traditionally considered Saltasauridae and a clade containing taxa normally found in Aeolosaurini and Lognkosauria.^[1] Unpublished cladistic results by Mocho et al. (2019) recover Lohuecotitan and Paludititan as members of Lirainosaurinae, and place Lirainosaurinae as an early-diverging branch of Saltasauridae.^[3]

Lithostrotia	Malawisaurus Paludititan Lohuecotitan Epachthosaurus Opisthocoelicaudiinae Alamosaurus Opisthocoelicaudia Saltasaurinae Neuquensaurus Rocasaurus Saltasaurus Lirainosaurinae Lirainosaurus Atsinganosaurus Ampelosaurus Bonatitan Rapetosaurus Nemegtosaurus Aeolosaurinae Gondwanatitan Aeolosaurus Rinconsauria Rinconsaurus Muyelensaurus Bonitasaura Lognkosauria Mendozasaurus Futalognkosaurus