List of erinaceids

Erinaceidae is a family of small mammals in the order Eulipotyphla. A member of this family is called an erinaceid, and the family includes hedgehogs and gymnures. Erinaceidae is one of four families in the order Eulipotyphla.^[1] They are found in Africa, Europe, and Asia, primarily in forests, shrublands, savannas, and grasslands, though some species can also be found in deserts, rocky areas, or caves. They range in size from the gymnures in the Hylomys genus, at 9 cm (4 in) plus a 1 cm (0.4 in) tail, to the moonrat, at 46 cm (18 in) plus a 30 cm (12 in) tail. Erinaceids are omnivorous and primarily eat insects and small vertebrates such as lizards, though they also consume plants, eggs, and fungi. Hedgehogs all have spines on their backs, while gymnures have fur. No erinaceids have population estimates, but the Hainan gymnure and Dinagat gymnure are categorized as endangered species.

European hedgehog (Erinaceus europaeus)

The twenty-four extant species of Erinaceidae are divided into two subfamilies: Erinaceinae, containing sixteen hedgehog species in five genera, and Galericinae, containing eight gymnure species in five genera. A few extinct prehistoric Erinaceidae species have been discovered, though due to ongoing research and discoveries the exact number and categorization is not fixed.^[2]

Conventions

IUCN Red List categories

Conservation status
EX	Extinct (0 species)
EW	Extinct in the wild (0 species)
CR	Critically endangered (0 species)
EN	Endangered (2 species)
VU	Vulnerable (1 species)
NT	Near threatened (1 species)
LC	Least concern (19 species)
Other categories
DD	Data deficient (1 species)
NE	Not evaluated (0 species)

The author citation for the species or genus is given after the scientific name; parentheses around the author citation indicate that this was not the original taxonomic placement. Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species. Range maps are provided wherever possible; if a range map is not available, a description of the erinaceid's range is provided. Ranges are based on the IUCN Red List for that species unless otherwise noted.

Classification

The family Erinaceidae consists of two subfamilies: Erinaceinae, containing sixteen hedgehog species in five genera, and Galericinae, containing eight gymnure species in five genera.

Family Erinaceidae

• Subfamily Erinaceinae
  • Genus Atelerix (African hedgehogs): four species
  • Genus Erinaceus (woodland hedgehogs): four species
  • Genus Hemiechinus (long-eared hedgehogs): two species
  • Genus Mesechinus (steppe hedgehogs): two species
  • Genus Paraechinus (desert hedgehogs): four species
• Subfamily Galericinae
  • Genus Echinosorex (moonrat): one species
  • Genus Hylomys (gymnures): three species
  • Genus Neohylomys (Hainan gymnure): one species
  • Genus Neotetracus (shrew gymnure): one species
  • Genus Podogymnura (Philippine gymnures): two species

Erinaceidae[3] Erinaceinae Mesechinus Hemiechinus Paraechinus Atelerix Erinaceus Galericinae Echinosorex Podogymnura Hylomys Neohylomys Neotetracus

Erinaceids

The following classification is based on the taxonomy described by the reference work Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis, as supported by both the IUCN and the American Society of Mammalogists.^[1] The range shown is only for wild populations; domesticated hedgehogs, which are usually four-toed hedgehogs, North African hedgehogs, or hybrids of the two, may live outside these areas.

Subfamily Erinaceinae

Main article: Erinaceinae

Genus Atelerix – Pomel, 1848 – four species

Common name	Scientific name and subspecies	Range	Size and ecology	IUCN status and estimated population
Four-toed hedgehog	A. albiventris (Wagner, 1841)	Western, central, and eastern Africa	Size: 17–23 cm (7–9 in) long[4] Habitat: Savanna and grassland[5] Diet: Invertebrates, as well as plants and small vertebrates[4]	LC  Unknown [5]
North African hedgehog	A. algirus (Lereboullet, 1842) Three subspecies A. a. algirus A. a. girbanensis A. a. vagans	Northern Africa	Size: 20–25 cm (8–10 in) long[6] Habitat: Forest, shrubland, and grassland[7] Diet: Omnivorous, including arthropods, small vertebrates, carrion, and fungi[6]	LC  Unknown [7]
Somali hedgehog	A. sclateri Anderson, 1895	Somalia	Size: 20–27 cm (8–11 in) long, plus 1–2 cm (0.4–0.8 in) tail[8] Habitat: Grassland and savanna[9] Diet: Believed to be omnivorous; primarily insects[8]	LC  Unknown [9]
Southern African hedgehog	A. frontalis (Smith, 1831)	Southern Africa	Size: 15–20 cm (6–8 in) long, plus 2 cm (1 in) tail[10] Habitat: Savanna, shrubland, and grassland[11] Diet: Omnivorous; primarily insects, as well as carrion, vegetables, fungi, and small vertebrates[10]	LC  Unknown [11]

Genus Erinaceus – Linnaeus, 1758 – four species

Common name	Scientific name and subspecies	Range	Size and ecology	IUCN status and estimated population
Amur hedgehog	E. amurensis Schrenk, 1859	Eastern Asia	Size: 15–29 cm (6–11 in) long, plus 1–5 cm (0.4–2.0 in) tail[12] Habitat: Forest and shrubland[13] Diet: Earthworms and other ground invertebrates, as well as small vertebrates and fruit[13]	LC  Unknown [13]
European hedgehog	E. europaeus Linnaeus, 1758	Europe, northwestern Asia, and New Zealand	Size: 13–27 cm (5–11 in) long, plus 2 cm (1 in) tail[14] Habitat: Forest and grassland[15] Diet: Omnivorous; primarily insects, as well as other invertebrates, eggs, small vertebrates, and carrion[14]	NT  Unknown [15]
Northern white-breasted hedgehog	E. roumanicus Barrett-Hamilton, 1900	Eastern Europe and western Asia (in blue)	Size: 13–30 cm (5–12 in) long, plus 1–5 cm (0.4–2.0 in) tail[16] Habitat: Forest and shrubland[17] Diet: Omnivorous; primarily insects, as well as other invertebrates, eggs, small vertebrates, and carrion[18]	LC  Unknown [17]
Southern white-breasted hedgehog	E. concolor Martin, 1838	Western Asia (in orange)	Size: 13–30 cm (5–12 in) long, plus 1–5 cm (0.4–2.0 in) tail[16] Habitat: Forest and shrubland[19] Diet: Omnivorous; primarily insects, as well as other invertebrates, eggs, small vertebrates, and carrion[18]	LC  Unknown [19]

Genus Hemiechinus – Fitzinger, 1866 – two species

Common name	Scientific name and subspecies	Range	Size and ecology	IUCN status and estimated population
Indian long-eared hedgehog	H. collaris (Gray, 1830)	Western India and Pakistan	Size: 15–28 cm (6–11 in) long, plus 1–6 cm (0.4–2.4 in) tail[20] Habitat: Shrubland, grassland, and desert[21] Diet: Omnivorous; primarily invertebrates and insects, as well as small vertebrates, eggs, carrion, fruit, and seeds[20]	LC  Unknown [21]
Long-eared hedgehog	H. auritus (Gmelin, 1770)	Western and central Asia	Size: 12–27 cm (5–11 in) long, plus 1–5 cm (0.4–2.0 in) tail[22] Habitat: Shrubland and desert[23] Diet: Omnivorous; primarily invertebrates and insects, as well as eggs, fruit, vegetables, and small vertebrates[22]	LC  Unknown [23]

Genus Mesechinus – Ognew, 1951 – two species

Common name	Scientific name and subspecies	Range	Size and ecology	IUCN status and estimated population
Daurian hedgehog	M. dauuricus (Sundevall, 1842)	East-central Asia	Size: About 24 cm (9 in) long, plus 3 cm (1 in) tail[20] Habitat: Forest and grassland[24] Diet: Beetles and other invertebrates, as well as small reptiles, bird eggs and nestlings, rodents, and carrion[24]	LC  Unknown [24]
Hugh's hedgehog	M. hughi (Thomas, 1908)	Central China	Size: About 24 cm (9 in) long, plus 3 cm (1 in) tail[20] Habitat: Grassland[25] Diet: Omnivorous; primarily invertebrates and insects, as well as small vertebrates, eggs, carrion, fruit, and seeds[20]	LC  Unknown [25]

Genus Paraechinus – Trouessart, 1879 – four species

Common name	Scientific name and subspecies	Range	Size and ecology	IUCN status and estimated population
Bare-bellied hedgehog	P. nudiventris (Horsfield, 1851)	Southern India	Size: 14–28 cm (6–11 in) long, plus 1–4 cm (0.4–1.6 in) tail[26] Habitat: Forest and shrubland[27] Diet: Insects, as well as small vertebrates, eggs, and scorpions[20]	LC  Unknown [27]
Brandt's hedgehog	P. hypomelas (Brandt, 1836)	Western Asia	Size: 14–28 cm (6–11 in) long, plus 1–4 cm (0.4–1.6 in) tail[26] Habitat: Savanna, shrubland, grassland, and desert[28] Diet: Insects, as well as small vertebrates, eggs, and scorpions[20]	LC  Unknown [28]
Desert hedgehog	P. aethiopicus (Ehrenberg, 1832)	Northern Africa and Arabian Peninsula	Size: 14–23 cm (6–9 in) long[29] Habitat: Desert, inland wetlands, grassland, shrubland, and savanna[30] Diet: Insects, as well as other invertebrates, amphibians, reptiles, and eggs[29]	LC  Unknown [30]
Indian hedgehog	P. micropus (Blyth, 1846)	Western India and Pakistan	Size: 14–28 cm (6–11 in) long, plus 1–4 cm (0.4–1.6 in) tail[31] Habitat: Shrubland and desert[32] Diet: Insects, as well as other invertebrates, small vertebrates, and eggs[31]	LC  Unknown [32]

Subfamily Galericinae

Main article: Galericinae

Genus Echinosorex – Blainville, 1836 – one species

Common name	Scientific name and subspecies	Range	Size and ecology	IUCN status and estimated population
Moonrat	E. gymnura (Raffles, 1822) Two subspecies E. g. alba E. g. gymnura	Southeast Asia	Size: 26–46 cm (10–18 in) long, plus 16–30 cm (6–12 in) tail[33] Habitat: Forest[34] Diet: Invertebrates, as well as frogs, fish, and fruit[35]	LC  Unknown [34]

Genus Hylomys – Müller, 1839 – three species

Common name	Scientific name and subspecies	Range	Size and ecology	IUCN status and estimated population
Dwarf gymnure	H. parvus Robinson & Kloss, 1916	Sumatra island in Indonesia	Size: 9–15 cm (4–6 in) long, plus 1–3 cm (0.4–1.2 in) tail[36] Habitat: Forest[37] Diet: Invertebrates, as well as fruit[36]	VU  Unknown [37]
Long-eared gymnure	H. megalotis Jenkins & Robinson, 2002	Laos	Size: 9–15 cm (4–6 in) long, plus 1–3 cm (0.4–1.2 in) tail[36] Habitat: Forest, shrubland, and rocky areas[38] Diet: Invertebrates, as well as fruit[36]	DD  Unknown [38]
Short-tailed gymnure	H. suillus Müller, 1840 Seven subspecies H. s. dorsalis (Bornean short-tailed gymnure) H. s. maxi (Max's short-tailed gymnure) H. s. microtinus H. s. pegunensis (Northern short-tailed gymnure) H. s. siamensis H. s. suillus (Javan short-tailed gymnure) H. s. tionis	Southeastern Asia	Size: 9–15 cm (4–6 in) long, plus 1–3 cm (0.4–1.2 in) tail[36] Habitat: Forest and shrubland[39] Diet: Invertebrates, as well as fruit[39]	LC  Unknown [39]

Genus Neohylomys – Shaw & Wong, 1959 – one species

Common name	Scientific name and subspecies	Range	Size and ecology	IUCN status and estimated population
Hainan gymnure	N. hainanensis Shaw & Wong, 1959	Hainan island, China	Size: 12–15 cm (5–6 in) long, plus 3–5 cm (1–2 in) tail[40] Habitat: Forest and caves[41] Diet: Insects, worms, and plants[40]	EN  Unknown [41]

Genus Neotetracus – Trouessart, 1909 – one species

Common name	Scientific name and subspecies	Range	Size and ecology	IUCN status and estimated population
Shrew gymnure	N. sinensis Trouessart, 1909	Southern China	Size: 10–13 cm (4–5 in) long, plus 4–7 cm (2–3 in) tail[40] Habitat: Forest[42] Diet: Insects, worms, and plants[40]	LC  Unknown [42]

Genus Podogymnura – Mearns, 1905 – two species

Common name	Scientific name and subspecies	Range	Size and ecology	IUCN status and estimated population
Dinagat gymnure	P. aureospinula Heaney & Morgan, 1982	Philippines	Size: 19–22 cm (7–9 in) long, plus 5–8 cm (2–3 in) tail[16] Habitat: Forest[43] Diet: Insects, worms, and birds[16]	EN  Unknown [43]
Mindanao gymnure	P. truei Mearns, 1905	Philippines	Size: 13–15 cm (5–6 in) long, plus 4–7 cm (2–3 in) tail[44] Habitat: Forest[45] Diet: Insects, worms, and carrion[44]	LC  Unknown [45]