Lithognathus

Lithognathus is a genus of marine ray-finned fish belonging to the family Sparidae, which includes the seabreams and porgies. Species in this genus are given the common name of steenbras. The genus is found in the Eastern Atlantic Ocean from southwestern Europe to South Africa and into the southwestern Indian Ocean.

Lithognathus Temporal range: Miocene to Present[1]
Lithognathus mormyrus
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Actinopterygii
Order:	Acanthuriformes
Family:	Sparidae
Genus:	Lithognathus Swainson, 1839[2]
Type species
Lithognathus capensis Swainson, 1839[3]
Species
See text
Synonyms[3]
Pagrichthys Bleeker, 1859

Taxonomy

Lithognathus was first proposed as taxon in 1839 by the English zoologist William Swainson, Swainson named it as a monotypic subgenus of Pagellus with Pagellus (Lithognathis) capensis as its only species and, therefore, its type species.^[3] Swainson's name is now understood to be a junior synonym of Pagrus lithognathus which had been described in 1829 by Georges Cuvier, with its type locality given as the Cape of Good Hope.^[4] The genus Lithognathus is placed in the family Sparidae within the order Spariformes by the 5th edition of Fishes of the World.^[5] Some authorities classify this genus in the subfamily Pagellinae,^[6] but the 5th edition of Fishes of the World does not recognise subfamilies within the Sparidae.^[5]

Etymology

Lithognathus means "stone jaw", Swainson described the maxillaries as "thick, enlarged, and as hard as stone". It is not a tautonym as Swainson unnecessarily renamed Cuvier's Pagellus lithognathus as L. lithognathus.^[7]

Species

The World Register of Marine Species lists the following four species :^[8]

• Lithognathus aureti Smith, 1962 (West coast seabream)
• Lithognathus lithognathus (Cuvier, 1829) (White steenbras)
• Lithognathus mormyrus (Linnaeus, 1758) (Sand steenbras)
• Lithognathus olivieri Penrith & Penrith, 1969 (Steenbras)

Characteristics

Lithognathus steenbras breams are characterised by an oblong compressed, body with a long snout. The tips of the pectoral fins extend almost as far as the origin of the anal fin. The teeth in the front of the jaws are small and arranged in bands with between 3 and 6 rows of molar-like teeth in the upper jawand between 2 and 4 rows in the lower jaw. In adults the maxillasoes not extend as far as the anterior edge of the Orbit (anatomy), The scales on the head reach as afr as the posterior edge of the orbit.^[9] The largest species in the genus is the white steenbras with a maximum published total length of 200 cm (79 in) while the smallest is the L. olivieri.^[8]

Distribution

Lithognathus steenbras breams occur in the Eastern Atlantic Ocean from southwestern Europe to South Africa and into the southwestern Indian Ocean.^[8]