Macrobiotidae

Macrobiotidae is a family of tardigrades.

Macrobiotidae
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Tardigrada
Class:	Eutardigrada
Order:	Parachela
Superfamily:	Macrobiotoidea
Family:	Macrobiotidae Thulin, 1928[1]
Type genus
Macrobiotus C.A.S. Schultze, 1834

Phylogeny

External phylogeny

According to a 2018 report using multiple morphological and molecular studies, the Macrobiotoidea are sister to the Hypsibioidea; both are eutardigrades.^[2] The genera Adorybiotus and Richtersius were transferred from Macrobiotidae to a new family, Richtersiidae, in 2016.^[3][4]

Eutardigrada	Apochela Milnesiidae Parachela Isohypsibiodea Macrobiotoidea Macrobiotidae Richtersiidae[3] Hypsibioidea

Internal phylogeny

A 2021 phylogenetic analysis found that the genus Macrobiotus is monophyletic (a clade) but that the former genus Xerobiotus is wholly contained within Macrobiotus. A new genus, Sisubiotus, was created. The genus "Minibiotus" was seen to be paraphyletic with respect to Tenuibiotus and Paramacrobiotus.^[5]

Macrobiotidae	Macrobiotus Mesobiotus Sisubiotus "Minibiotus" (part) "Minibiotus" (part) Tenuibiotus Paramacrobiotus

Genera

As of 2023^[update], the family Macrobiotidae consists of the following genera:^[3]

• Biserovus Guidetti & Pilato, 2003
• Calcarobiotus Dastych, 1993
  • C. (Calcarobiotus) Dastych, 1993
  • C. (Discrepunguis) Guidetti & Bertolani, 2001
• Famelobiotus Pilato, Binda & Lisi, 2004
• Insuetifurca Guidetti & Pilato, 2003
• Macrobiotus C.A.S. Schultze, 1834
• Mesobiotus Vecchi et al., 2016
• Minibiotus R.O. Schuster, 1980
• Minilentus Guidetti & Pilato, 2003
• Paramacrobiotus Guidetti et al., 2009^[6][7][8]
• Pseudodiphascon Ramazzotti, 1965 (genus dubium)
• Pseudohexapodibius Bertolani & Biserov, 1996
• Schusterius Kaczmarek & Michalczyk, 2006
• Sisubiotus Stec, Vecchi, Calhim & Michalczyk, 2021
• Tenuibiotus Pilato & Lisi, 2011
• Xerobiotus Bertolani & Biserov, 1996