Papiliolebias

Papiliolebias is a genus of fish in the family Rivulidae. These annual killifish are endemic to seasonal pools in the Paraguay and upper Madeira river basins in northwestern Argentina, central and southeastern Bolivia, and western Paraguay.^[1][2][3]

Papiliolebias
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Actinopterygii
Order:	Cyprinodontiformes
Family:	Rivulidae
Genus:	Papiliolebias W. J. E. M. Costa, 1998
Type species
Plesiolebias bitteri W. J. E. M. Costa, 1989

They are small fish, up to 5 cm (2.0 in) in total length.^[4] As typical of killifish, males are more colorful than females.^[1][2][3]

Species

Papiliolebias and the closely related Maratecoara, Pituna, Plesiolebias and Stenolebias form a clade, Plesiolebiasini.^[5]

There are currently 5 recognized species in Papiliolebias:^[4]

• Papiliolebias ashleyae D. T. B. Nielsen & Brousseau, 2014^[1]
• Papiliolebias bitteri (W. J. E. M. Costa, 1989)
• Papiliolebias francescae Valdesalici & Brousseau, 2014^[2]
• Papiliolebias habluetzeli Valdesalici, D. T. B. Nielsen, Brousseau & Phunkner, 2016^[3]
• Papiliolebias hatinne Azpelicueta, Butí & G. B. García, 2009