Patrick J. Keeling

Patrick John Keeling is a biologist and professor in the Department of Botany at the University of British Columbia.^[6][3][7] His research investigates the phylogeny, genomics and molecular evolution of protists and his work has led to numerous advances in assembling the eukaryotic tree of life. He has also identified several cases of horizontal gene transfer.^[5][8][9]

Patrick Keeling
Born	Patrick John Keeling 1969 (age 55–56)
Alma mater	University of Western Ontario (BSc) Dalhousie University (PhD)
Awards	Guggenheim Fellowship (2011)[1] Gilbert Morgan Smith Medal (2021)[2]
Scientific career
Fields	Evolution Microbiology Protists[3]
Institutions	University of British Columbia University of Melbourne Indiana University Bloomington
Thesis	Studies on the prokaryote-eukaryote transition (1996)
Doctoral advisor	Ford Doolittle[4]
Other academic advisors	Jeffrey D. Palmer (postdoc)[5]
Website	www.botany.ubc.ca/people/patrick-keeling