Pea enation mosaic virus

Pea enation mosaic virus (PEMV) is two plant pathogenic viruses. The two viruses, once classified as one, are now categorised as two separate, mutualistic viruses:^[1]

• Pea enation mosaic virus 1 is an Enamovirus
• Pea enation mosaic virus 2 is an Umbravirus

Pea enation mosaic virus 1
Virus classification
(unranked):	Virus
Realm:	Riboviria
Kingdom:	Orthornavirae
Phylum:	Pisuviricota
Class:	Pisoniviricetes
Order:	Sobelivirales
Family:	Solemoviridae
Genus:	Enamovirus
Species:	Enamovirus PEMV

Pea enation mosaic virus 2
Virus classification
(unranked):	Virus
Realm:	Riboviria
Kingdom:	Orthornavirae
Phylum:	Kitrinoviricota
Class:	Tolucaviricetes
Order:	Tolivirales
Family:	Tombusviridae
Genus:	Umbravirus
Species:	Umbravirus pisi

They are spread by green- or pea aphids and affects such legumes as pea,^[2] alfalfa, broadbean or sweet pea mostly in temperate regions.^[1]

Symptoms include chlorotic, translucent or necrotic lesions, malformation of leaves and stipules, and plant distortion. However, the most characteristic symptom is the formation of enations on the abaxial, i.e. downy, leaf side. Enations are derived from the cells of vascular bundles undergoing hyperplasia.^[3]