Physalaemus

Physalaemus is a large genus of leptodactylid frogs. These frogs, sometimes known as dwarf frogs or foam frogs, are found in South America.^[1] It is very similar to Leptodactylus, a close relative, and indeed the recently described Leptodactylus lauramiriamae is in some aspects intermediate between them.^[2]

Physalaemus
Physalaemus cuvieri
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Amphibia
Order:	Anura
Family:	Leptodactylidae
Subfamily:	Leiuperinae
Genus:	Physalaemus Fitzinger, 1826
Type species
Physalaemus cuvieri Fitzinger, 1826
Diversity
About 48 species (see text)
Synonyms
Paludicola Wagler, 1830 Liuperus Cope, 1861 "1860" Gomphobates Reinhardt and Lütken, 1862 "1861" Eupemphix Steindachner, 1863 Nattereria Steindachner, 1864

Physalaemus marmoratus

Physalaemus maculiventris

Physalaemus olfersii

Species

There are 50^[1] or 49 species (AmphibiaWeb lists Physalaemus nattereri as Eupemphix nattereri ):^[3]

• Physalaemus aguirrei Bokermann, 1966
• Physalaemus albifrons (Spix, 1824)
• Physalaemus albonotatus (Steindachner, 1864)
• Physalaemus angrensis Weber, Gonzaga, and Carvalho-e-Silva, 2005
• Physalaemus araxa Leal, Zornosa-Torres, Augusto-Alves, Dena, Pezzuti, Leite, Lourenço, Garcia, and Toledo, 2021
• Physalaemus atim Brasileiro and Haddad, 2015
• Physalaemus atlanticus Haddad and Sazima, 2004
• Physalaemus barrioi Bokermann, 1967
• Physalaemus biligonigerus (Cope, 1861)
• Physalaemus bokermanni Cardoso and Haddad, 1985
• Physalaemus caete Pombal and Madureira, 1997
• Physalaemus camacan Pimenta, Cruz & Silvano, 2005
• Physalaemus carrizorum Cardozo and Pereyra, 2018
• Physalaemus centralis Bokermann, 1962
• Physalaemus cicada Bokermann, 1966
• Physalaemus claptoni Leal, Leite, Costa, Nascimento, Lourenço, and Garcia, 2020
• Physalaemus crombiei Heyer and Wolf, 1989
• Physalaemus cuqui Lobo, 1993
• Physalaemus cuvieri Fitzinger, 1826
• Physalaemus deimaticus Sazima and Caramaschi, 1988
• Physalaemus ephippifer (Steindachner, 1864)
• Physalaemus erikae Cruz and Pimenta, 2004
• Physalaemus erythros Caramaschi, Feio, and Guimarães, 2003
• Physalaemus evangelistai Bokermann, 1967
• Physalaemus feioi Cassini, Cruz, and Caramaschi, 2010
• Physalaemus fernandezae (Müller, 1926)
• Physalaemus fischeri (Boulenger, 1890)
• Physalaemus gracilis (Boulenger, 1883)
• Physalaemus henselii (Peters, 1872)
• Physalaemus insperatus Cruz, Cassini, and Caramaschi, 2008
• Physalaemus irroratus Cruz, Nascimento, and Feio, 2007
• Physalaemus jordanensis Bokermann, 1967
• Physalaemus kroyeri (Reinhardt and Lütken, 1862)
• Physalaemus lateristriga (Steindachner, 1864)
• Physalaemus lisei Braun and Braun, 1977
• Physalaemus maculiventris (Lutz, 1925)
• Physalaemus marmoratus (Reinhardt and Lütken, 1862)
• Physalaemus maximus Feio, Pombal, and Caramaschi, 1999
• Physalaemus moreirae (Miranda-Ribeiro, 1937)
• Physalaemus nanus (Boulenger, 1888)
• Physalaemus nattereri (Steindachner, 1863)
• Physalaemus obtectus Bokermann, 1966
• Physalaemus olfersii (Lichtenstein and Martens, 1856)
• Physalaemus orophilus Cassini, Cruz, and Caramaschi, 2010
• Physalaemus riograndensis Milstead, 1960
• Physalaemus rupestris Caramaschi, Carcerelli, and Feio, 1991
• Physalaemus santafecinus Barrio, 1965
• Physalaemus signifer (Girard, 1853)
• Physalaemus soaresi Izecksohn, 1965
• Physalaemus spiniger (Miranda-Ribeiro, 1926)