Pronephrium

Pronephrium is a genus of ferns in the family Thelypteridaceae, subfamily Thelypteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I).^[1] Other sources sink Pronephrium into a very broadly defined genus Thelypteris.^[3] Some species were split off into the genera Abacopteris, Grypothrix and Menisciopsis in 2021 as a result of a phylogenetic study of the family Thelypteridaceae.^[4][5]

Pronephrium
Pronephrium hewittii in Malaysia
Scientific classification
Kingdom:	Plantae
Clade:	Tracheophytes
Division:	Polypodiophyta
Class:	Polypodiopsida
Order:	Polypodiales
Suborder:	Aspleniineae
Family:	Thelypteridaceae
Subfamily:	Thelypteridoideae
Genus:	Pronephrium C.Presl[1]
Species
See text.
Synonyms[2]
Dimorphopteris Tagawa & K.Iwats. Haplodictyum C.Presl Nannothelypteris Holttum Pronephrium subg. Menisciopsis Holttum Thelypteris subg. Dimorphopteris K.Iwats. Thelypteris subg. Haplodictyum K.Iwats.

The genus was first described in 1851 by Carl Borivoj Presl.^[6][7]

Species

As of June 2022^[update], the Checklist of Ferns and Lycophytes of the World accepted the following species and hybrids:^[2]

• Pronephrium affine (Blume) C.Presl
• Pronephrium amboinense (Willd.) Holttum
• Pronephrium amphitrichum Holttum
• Pronephrium aoristisorum (Harr.) S.E.Fawc. & A.R.Sm.
• Pronephrium aquatiloides (Copel.) Holttum
• Pronephrium borneense (Hook.) Holttum
• Pronephrium camarinense (Holttum) S.E.Fawc. & A.R.Sm.
• Pronephrium celebicum (Baker) Holttum
• Pronephrium clemensiae (Copel.) Holttum
• Pronephrium euryphyllum (Rosenst.) Holttum
• Pronephrium exsculptum (Baker) Holttum
• Pronephrium fidelei Rakotondr.
• Pronephrium firmulum (Baker) Holttum
• Pronephrium giluwense Holttum
• Pronephrium granulosum (C.Presl) Holttum
• Pronephrium hewittii (Copel.) Holttum
• Pronephrium hosei (Baker) Holttum
• Pronephrium inaequilobatum (Holttum) S.E.Fawc. & A.R.Sm.
• Pronephrium kjellbergii Holttum
• Pronephrium lineatum (Blume) C.Presl
• Pronephrium manuselense M.Kato
• Pronephrium marojejyensis Rakotondr.
• Pronephrium menisciicarpon (Blume) Holttum
• Pronephrium merrillii (Christ) Holttum
• Pronephrium millarae Holttum
• Pronephrium minahassae Holttum
• Pronephrium moniliforme (Tagawa & K.Iwats.) Holttum
• Pronephrium murkelense (M.Kato) S.E.Fawc. & A.R.Sm.
• Pronephrium nervosum (Fee) S.E.Fawc. & A.R.Sm.
• Pronephrium palauense (Hosok.) Holttum
• Pronephrium peltatum (Alderw.) Holttum
• Pronephrium peramalense Holttum
• Pronephrium philippinum (C.Presl) S.E.Fawc. & A.R.Sm.
• Pronephrium rhombeum (Christ) Holttum
• Pronephrium samarense (Copel.) Holttum
• Pronephrium simillimum (C.Chr.) Holttum
• Pronephrium solsonicum Holttum
• Pronephrium thysanoides Holttum
• Pronephrium trachyphyllum Holttum
• Pronephrium xiphioides (Christ) Holttum

Hybrids

• Pronephrium × interruptum Holttum
• Pronephrium × liukiuense (Christ) Nakaike
• Pronephrium × occultum (C.Hope) comb.ined.