Pusula

This article is about the snail genus. For the municipality of Finland, see Nummi-Pusula.

Pusula is a genus of small sea snails, marine gastropod mollusks in the family Triviidae, the false cowries or trivias.^[1][2]

Pusula
A shell of Pusula radians from Nicaragua on display at the Museo Civico di Storia Naturale di Milano
Scientific classification
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
(unranked):	clade Caenogastropoda clade Hypsogastropoda clade Littorinimorpha
Superfamily:	Velutinoidea
Family:	Triviidae
Subfamily:	Triviinae
Genus:	Pusula Jousseaume, 1884
Type species
Cypraea radians Lamarck, 1810
Synonyms[1]
Discotrivia Cate, 1979

Species

Species within the genus Pusula include:

• † Pusula alienigena Schilder, 1928
• Pusula bessei Petuch, 2013
• Pusula cimex (Sowerby, 1870)
• Pusula costipunctata (Sowerby, 1870)
• † Pusula dadeensis Petuch, 1994
• † Pusula dalli (Petuch, 1994)
• † Pusula densistans Fehse & Grego, 2014^[2]
• Pusula elsiae (Howard & Sphon, 1960) (nomen dubium)
• Pusula garciai Fehse & Grego, 2014^[2]
• † Pusula guppyi Schilder, 1939
• Pusula hybrida (Schilder, 1931)
• Pusula labiosa (Gaskoin, 1836)
• Pusula lindajoyceae Petuch, 1994
• † Pusula miamiensis Petuch, 1991
• † Pusula orientalis Schilder, 1939
• Pusula pediculus (Linnaeus, 1758)^[3]
• † Pusula permagna (Johnson, 1910)
• † Pusula platyventer Fehse & Grego, 2014 ^[2]
• Pusula pullata (Sowerby, 1870)
• Pusula radians (Lamarck, 1810)
• Pusula solandri (Sowerby, 1832)
• † Pusula subpediculus (Sacco, 1894)

Species brought into synonymy

• † Pusula andersoni Petuch & Drohlshagen, 2011: synonym of † Pusula lindajoyceae Petuch, 1994
• Pusula bermontiana (Petuch, 1994): synonym of Pusula pediculus (Linnaeus, 1758)
• Pusula californiana (Gray, 1827): synonym of Pseudopusula californiana (Gray, 1827)
• Pusula californica (Sowerby, 1832): synonym of Pseudopusula californiana (Gray, 1827)
• Pusula calusa Petuch & Drolshagen, 2011: synonym of † Pusula dalli (Petuch, 1994)
• Pusula campus Cate, 1979: synonym of Niveria campus (Cate, 1979)
• Pusula candidula (Gaskoin, 1835):^[4] synonym of Trivia candidula (Gaskoin, 1836)
• Pusula carabus Cate, 1979: synonym of Niveria carabus (Cate, 1979)
• Pusula cherobia Cate, 1979: synonym of Niveria cherobia (Cate, 1979)
• Pusula crovoae Olsson, 1967: synonym of † Pusula orientalis Schilder, 1939
• Pusula depauperata (Sowerby, 1832): synonym of Pseudopusula depauperata (Sowerby, 1832)
• Pusula janae Lorenz, 2001:^[5] synonym of Dolichupis janae (Lorenz, 2001)
• Pusula juyingae Petuch & Sargent, 2011: synonym of Pusula labiosa (Gaskoin, 1836)
• Pusula leucosphaera Schilder, 1931: synonym of Dolichupis leucosphaera (Schilder, 1931)
• Pusula loochooensis Cate, 1979: synonym of Pseudopusula californiana (Gray, 1827)
• Pusula macaeica Fehse & Grego, 2005:^[6] synonym of Niveria macaeica (Fehse & Grego, 2005)
• Pusula maltbiana (Schwengel & McGinty, 1942):^[7] synonym of Niveria maltbiana (Schwengel & McGinty, 1942)
• Pusula olssoni Petuch & Drolshagen, 2011 †: synonym of † Pusula orientalis Schilder, 1939
• Pusula pacei (Petuch, 1987): synonym of Pusula cimex (Sowerby, 1870)
• Pusula padreserrai Cate, 1979: synonym of Pusula solandri (Sowerby, 1832)
• Pusula pygmaea Schilder, 1931: synonym of Cleotrivia pygmaea (Schilder, 1931)
• Pusula rubinicolor (Gaskoin, 1836): synonym of Dolichupis rubinicolor (Gaskoin, 1836)