Quadrula

Quadrula is a genus of freshwater mussels, aquatic bivalve mollusks in the family Unionidae native to rivers of the American Midwest and mid-south. All have thick nacreous shells with well-developed hinge teeth, many also with external shell sculpturing of nodules or lumps.

Quadrula
Quadrula fragosa
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Bivalvia
Order:	Unionida
Family:	Unionidae
Tribe:	Quadrulini
Genus:	Quadrula Rafinesque, 1820

Species within the genus Quadrula

In 2012, many species classified as Quadrula were moved to Rotundaria and Theliderma based on genetic and morphological evidence.^[1] Species currently in Quadrula now consist of:^[2]

Quadrula verrucosa

• Quadrula apiculata (Southern mapleleaf)
• Quadrula fragosa (Winged mapleleaf)
• Quadrula nobilis (Gulf mapleleaf)
• Quadrula quadrula (Mapleleaf)
• Quadrula rumphiana (Ridged mapleleaf)
• Quadrula verrucosa (Pistolgrip)