Rhysodinae

Rhysodinae is a subfamily (sometimes called wrinkled bark beetles) in the family Carabidae. There are 19 genera and at least 380 described species in Rhysodinae.^[1][2][3] The group of genera making up Rhysodinae had been treated as the family Rhysodidae in the past, and subsequent DNA analysis then placed it within Carabidae, where it was sometimes treated as the tribe Rhysodini,^[4][5] but the most recent analyses place it as a subfamily in a clade along with subfamilies Paussinae and Siagoninae, forming a sister to the remaining Carabidae.^[6]

Rhysodinae
Omoglymmius americanus
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Coleoptera
Suborder:	Adephaga
Family:	Carabidae
Subfamily:	Rhysodinae Laporte, 1840

Description

These beetles are elongate, in size ranging from 5–8 mm, and color ranging from a reddish brown to black. Both the thorax and the elytra are deeply grooved lengthwise, thus giving these beetles their common name. The head is also grooved, and posteriorly constricted into a short but visible "neck". The 11-segment antennae are short, resembling a string of beads, while the mandibles lack cutting edges and are thus nonfunctional. The front legs are short and strongly built.^{[citation needed]}

Adults and larvae live in moist rotten wood that is infested with slime moulds, which are believed to be their diet. Instead of using their mandibles to bite, they use the anterior edge of the mentum and swivel their heads to cut off pieces of food. Adults do not make burrows, instead just squeezing between the cell layers of the decomposed wood, generally leaving no visible trace of their passage, while larvae live in short tunnels.^{[citation needed]}

They occur on all continents with forested areas, the richest fauna being found in New Guinea, Indonesia, the Philippines, and northern South America.

Genera

Position in relation to the Carabidae according to Vasilikopoulos et al., 2021

Cicindelidae ⊞ Manticorini Megacephalini Collyridini Cicindelini Oxycheilini ⊟ Carabidae Rhysodinae Paussinae Siagoninae ⊞other subfamilies Carabinae Nebriinae Broscinae Elaphrinae Loricerinae Omophroninae Scaritinae Trechinae Brachininae Harpalinae ⊟

Tribe Clinidiini R.T. & J.R.Bell, 1978

Clinidium Kirby, 1830

Grouvellina R.T. & J.R.Bell, 1978

Rhyzodiastes Fairmaire, 1895

Tribe Dhysorini R.T. & J.R.Bell, 1978

Dhysores Grouvelle, 1903

Neodhysores R.T. & J.R.Bell, 1978

Tangarona R.T. & J.R.Bell, 1982

Tribe Leoglymmiini R.T. & J.R.Bell, 1978

Leoglymmius R.T. & J.R.Bell, 1978

Tribe Medisorini R.T. & J.R.Bell, 1987

Medisores R.T. & J.R.Bell, 1987

Tribe Omoglymmiini R.T. & J.R.Bell, 1978

Arrowina R.T. Bell & J.R. Bell, 1978

Omoglymmius Ganglbauer, 1891

Plesioglymmius R.T. & J.R.Bell, 1978

Shyrodes Grouvelle, 1903

Srimara R.T. & J.R.Bell, 1978

Xhosores R.T. & J.R.Bell, 1978

Yamatosa R.T. & J.R.Bell, 1979

Tribe Rhysodini Laporte, 1840

Kaveinga R.T. & J.R.Bell, 1978

Kupeus R.T. & J.R.Bell, 1982

Rhysodes Germar, 1822

Tribe Sloanoglymmiini R.T. & J.R.Bell, 1991

Sloanoglymmius R.T. & J.R.Bell, 1991