Staphyliniformia

Staphyliniformia is a large infraorder of beetles. It contains over 70,000 described species from all regions of the world. Most species occur in moist habitats - various kinds of rotting plant debris, fungi, dung, carrion, many live in fresh water.

Staphyliniformia Temporal range: Rhaetian–Recent PreꞒ Ꞓ O S D C P T J K Pg N
Devil's coach horse beetle, Ocypus olens
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Coleoptera
Suborder:	Polyphaga
Infraorder:	Staphyliniformia Lameere, 1900
Superfamilies
See text.

Characteristics

Most Staphyliniforms are small to average sized beetles. The diverse group has few clear apomorphies. They have primitively 11-segmented antennae, a constricted neck well behind the eyes. The pronotum has a well defined, large lateral edge. Larval legs are 5-segmented, the 10th abdominal segment is often with more-or-less fine or strong spines or hooks. Urogomphi (paired "horns" at posterior tip of abdomen of larvae and pupae) with basal articulation.^[1]

Systematics and evolution

Staphyliniformia belongs to the suborder Polyphaga and is usually given an infraorder or series rank. It contains three superfamilies:

• Histeroidea, including the clown beetles.
• Hydrophiloidea, including the water scavenger beetles
• Staphylinoidea, including antlike stone beetles, carrion beetles, and rove beetles.

Some recent studies also include the superfamily Scarabaeoidea (infraorder Scarabaeiformia), forming together the so-called Hydrophiloid lineage.^[2][3] A sister group relationship of Hydrophiloidea and Histeroidea is strongly supported.^[4]

The unambiguous fossil record dates back to Triassic, and an early Mesozoic origin of the group is probable.^[5]

See also

• List of subgroups of the order Coleoptera