Trigonochlamydidae

Trigonochlamydidae is a family of air-breathing land slugs, terrestrial gastropod molluscs in the clade Eupulmonata (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).

Trigonochlamydidae Temporal range: Recent[1] PreꞒ Ꞓ O S D C P T J K Pg N ↓
Parmacellilla filipowitschi
Scientific classification
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
Order:	Stylommatophora
Superfamily:	Parmacelloidea
Family:	Trigonochlamydidae Hesse, 1882
Subfamilies and genera[2]
Parmacellillinae P. Hesse, 1926 Parmacellilla Simroth, 1910 Trigonochlamydinae P. Hesse, 1882 Boreolestes Schileyko & Kijashko, 1999 Drilolestes Lindholm, 1925 Hyrcanolestes Simroth, 1901 Khostalestes Suvorov, 2003 Lesticulus Schileyko, 1988 Pseudomilax O. Boettger, 1881 Trigonochlamys O. Boettger, 1881 Troglolestes Liovushkin & Matiokin, 1965
Diversity
9 genera, 11 species

Distribution

The native distribution of Trigonochlamydidae includes Caucasus,^[1] Iran and Turkey.^[3]

Taxonomy

Previously, Trigonochlamydidae was placed in the superfamily Trigonochlamydoidea Hesse, 1882, in the subinfraorder Sigmurethra. This was the only family within that superfamily.^[4]

The family Trigonochlamydidae is now classified within the limacoid clade, which itself belongs to the clade Stylommatophora, within the clade Eupulmonata (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).

The following two subfamilies have been recognized in the taxonomy of Bouchet & Rocroi (2005):

• subfamily Trigonochlamydinae Hesse, 1882 – synonyms: Selenochlamydinae I. M. Likharev & Wiktor, 1980
• subfamily Parmacellillinae Hesse, 1926^[5]

Genera

There are nine^[3] genera in the family Trigonochlamydidae with a total of 11 species:

Subfamily Trigonochlamydinae

• Boreolestes Schileyko & Kijashko, 1999
  • Boreolestes likharevi Schileyko & Kijashko, 1999 – type species^[6]
  • Boreolestes sylvestris Kijashko in Schileyko & Kijashko, 1999^[6]
• Drilolestes Lindholm, 1925 – with only one species Drilolestes retowskii (O. Boettger, 1884)^[6]
• Hyrcanolestes Simroth, 1901 – with only one species Hyrcanolestes velitaris (Martens, 1880)^[6]
• Khostalestes Suvorov, 2003^[3] – with only one species Khostalestes kochetkovi Suvorov, 2003^[3][6]
• Lesticulus Schileyko, 1988 – with only one species Lesticulus nocturnus Schileyko, 1988^[6]
• Trigonochlamys O. Boettger, 1881 – type genus of the family Trigonochlamydidae, with only one species Trigonochlamys imitatrix O. Boettger, 1881^[6]
• Troglolestes Ljovushkin & Matiokin, 1965 – with only one species Troglolestes sokolovi Liovushkin & Matiokin, 1965^[6]

Subfamily Parmacellillinae

• Parmacellilla Simroth, 1910 – type genus of the subfamily Parmacellillinae, with only one species Parmacellilla filipowitschi Simroth, 1910^[7]

Synonym:

• Pseudomilax O. Boettger, 1881 is a synonym for various genera in Trigonochlamydidae.

Selenochlamys O. Boettger, 1883 has been removed from the Trigonochlamydidae to the Oxychilidae based on DNA evidence.^[8]

Cladogram

The following cladogram shows the phylogenic relationships of this family with the other families in the limacoid clade:^[9]

limacoid clade	Staffordioidea Staffordiidae Dyakioidea Dyakiidae Gastrodontoidea Pristilomatidae Chronidae Euconulidae Trochomorphidae Gastrodontidae Oxychilidae Parmacelloidea Trigonochlamydidae Parmacellidae Milacidae Zonitoidea Zonitidae Helicarionoidea Helicarionidae Ariophantidae Urocyclidae Limacoidea Vitrinidae Boettgerillidae Limacidae Agriolimacidae