Woodsia

This article is about the genus of plants. For the genus of fish, see Woodsia (fish).

Woodsia is a genus of ferns in the family Woodsiaceae. Species of Woodsia are commonly known as cliff ferns.^[4]: 224 In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), it was the only genus in the family Woodsiaceae.^[2] In 2020, Physematium was split off from Woodsia on the basis of molecular phylogenetic evidence.^[5] As of June 2023^[update], Plants of the World Online continued to treat Physematium as a synonym of Woodsia.^[6]

Woodsia
Woodsia alpina[1]
Scientific classification
Kingdom:	Plantae
Clade:	Tracheophytes
Division:	Polypodiophyta
Class:	Polypodiopsida
Order:	Polypodiales
Suborder:	Aspleniineae
Family:	Woodsiaceae Herter[2]
Genus:	Woodsia R.Br.[2][3]
Type species
Woodsia ilvensis (von Linné) Brown
Species
See text
Synonyms
Eriosoriopsis (Kitagawa 1935) Ching & Wu 1991 Trichocyclus Dulac 1867 non N.E. Br. 1922

Phylogeny

There are about 25–30 species of the genus Woodsia. As of June 2023^[update], World Ferns hosted at World Plants accepted the following species:^[7]

Phylogeny from Fern Tree of Life[8][9]

Other species include:

Woodsia (Eriosorus) W. lanosa Hook. W. okamotoi Tagawa W. cycloloba Hand.-Mazz. W. kungiana Li Bing Zhang, N.T.Lu & X.F.Gao (Woodsia) section W. guizhouensis P.S.Wang, Q.Luo & Li Bing Zhang W. rosthorniana Diels Eriosoriopsis section Woodsia glabella R.Br. (smooth cliff fern) Woodsia alpina (Bolton) Gray (alpine woodsia) W. shensiensis Ching Alpinae section W. andersonii (Bedd.) Christ Woodsia ilvensis (L.) R.Br. (oblong woodsia, rusty cliff fern) W. gorovoii Krestsch. & Shmakov W. macrochlaena Mett. ex Kuhn Woodsia section W. oblonga Ching & S.H.Wu W. subcordata Turcz. W. polystichoides D.C.Eaton W. subintermedia Tzvelev Subcordatae

W. asiatica Kiselev & Shmakov W. calcarea (Fomin) Shmakov W. cinnamomea Christ W. hancockii Baker W. macrospora C.Chr. & Maxon W. nikkoensis H.Ogura & Nakaike W. pilosa Ching †W. pseudomanchuriensis Akhmetiev W. pseudopolystichoides (Fomin) Kiselev & Shmakov W. pulchella Bertol. W. sinica Ching W. taigischensis (Stepanov) Kuznetsov W. taishanensis F.Z.Li & C.K.Ni

Some hybrids are also known:^[7]

• Woodsia × gracilis (G.Lawson) Butters = W. alpina × W. ilvensis
• Woodsia × tryonis B.Boivin = W. ilvensis × W. glabella

When the genus Physematium is accepted, × Woodsimatium abbeae is a hybrid between Woodsia ilvensis and Physematium scopulinum.^[10]