Zhuganpo Formation

The Zhuganpo Formation is a Triassic geologic unit found in southern China. It has historically been known as the Zhuganpo Member of the Falang Formation. A diverse fossil assemblage known as the Xingyi biota or Xingyi Fauna can be found in the upper part of the Zhuganpo Formation. Fossils of the Xingyi biota include articulated skeletons of marine reptiles, abundant fish, and a plentiful assortment of invertebrates indicating a Ladinian to Carnian age for the sediments of the formation.^[1][2]

Zhuganpo Formation
Stratigraphic range: Upper Ladinian - Lower Carnian
Type	Geological formation
Underlies	Xiaowa Formation
Overlies	Yangliujing Formation
Lithology
Primary	limestone, marl
Location
Region	Guizhou Province, Yunnan Province
Country	China
Extent	Yunnan–Guizhou Plateau

Paleobiota

Invertebrates

• Ammonites: Clionitites sp.,^[3] Detoniceras sp.,^[3] Haoceras xingyiense,^[3][4] Parasturia sp.,^[3] Protrachyceras sp.,^[1] Ptychites sp.,^[3] Sinomeginoceras (S. wangi, S. xingyiense),^[3] Trachyceras sp.,^[3][4] Xenoprotrachyceras cf. primum,^[3] Yangites densicostatus,^[3]
• Bivalves: Daonella sp.^[1]
• Conodonts: Gladigondolella malayensis,^[4][5] Metapolygnathus / Paragondolella / Quadralella (Q. aff. acuminatus, P. foliata, P. inclinata, Q. intermedius, Q. langdaiensis, P. maantangensis, P. navicula, M. nodosus, M. parafoliata, Q. aff. praelindae, P. polygnathiformis, Q. shijiangjunensis, Q. tadpole, Q. uniformis, Q. wanlanensis, Q. aff. wayaoensis, Q. yongueensis, Q. aff. zonneveldi)^[1][4][5]
• Crinoids: Traumatocrinus hsui^[1]
• Crustaceans ("shrimps"): Schimperella acanthocercus (Lophogastrida)^[6][1]

Fish

• Acrolepidae indet.^[1]
• Archaesemionotus sp.^[1]
• aff. Arctacanthus sp. (Chimaeriformes?)^[7]
• Asialepidotus shingyiensis (Ionoscopiformes)^[1][8]
• Birgeria liui^[1][9]
• Caturidae indet.^[1]
• Eosemionotus sp.^[1]
• Favusodus orientalis (Euselachii)^[7]
• Fuyuanichthys wangi (Ginglymodi)^[10]
• Fuyuanperleidus dengi^[1]
• Guizhouamia bellula^[1]
• Guizhoubrachysomus minor^[1]
• Guizhoucoelacanthus guanlingensis^[1]
• Guizhouniscus microlepidus^[1]
• Habroichthys sp.^[1]
• Keichouodus nimaiguensis (Euselachii)^[7]
• Luganoia fortuna (Luganoiidae)^[11]
• Malingichthys (M. nimaiguensis, M. wanfendlinensis; Pholidophoridae)^[12]
• Marcopoloichthys sp.^[1]
• Peltopleurus (P. orientalis, P. tyrannos)^[1][13]
• Peripeltopleurus sp.^[1] (Wushaichthyidae)^[14]
• Pholidophoridae indet.^[1]
• Potanichthys xingyiensis (Thoracopteridae)^[15][1]
• Rosaodus xingyiensis (Elasmobranchii)^[7]
• Saurichthys sp.^[1]
• Sinoeugnathus kueichowensis^[1]
• Wushaichthys exquisitus^[16] (Wushaichthyidae)^[14]
• Xingyia gracilis^[1]

Reptiles

Reptiles of the Zhuganpo Formation
Genus	Species	Notes	Images
Anshunsaurus	A. huangnihensis[17]	An askeptosauroid thalattosaur[18]	Diandongosuchus fuyuanensis Keichousaurus hui Guizhouichthyosaurus
	A. wushaensis[19]	An askeptosauroid thalattosaur[20]
Brevicaudosaurus	B. jiyangshanensis	A small nothosaur with an unusually short torso and tail[21]
Diandongosuchus[22]	D. fuyuanensis	A semiaquatic archosauriform, interpreted as a basal phytosaur[23]
Dingxiaosaurus	D. luyinensis	A pistosauroid of uncertain validity.[24] Previously believed to have been from the Yangliujing Formation.[1]
Fuyuansaurus[25]	F. acutirostris	A "protorosaur" archosauromorph,[25] possibly a tanystropheid[26][27]
Glyphoderma[28]	G. kangi	A placochelyid placodont
Guizhouichthyosaurus	G. sp.[2][29]	A large predatory merriamosaurian ichthyosaur
Keichousaurus	K. hui	A keichousaurid pachypleurosaur[30][31]
Lariosaurus	L. xingyiensis	A small nothosaur[32][33]
Litorosuchus[34]	L. somnii	A semiaquatic archosauriform related to Vancleavea campi
Macrocnemus	M. fuyuanensis	A small basal tanystropheid[35]
Nothosaurus	N. youngi	A large nothosaur[36]
Qianichthyosaurus	Q. xingyiensis[37]	A small toretocnemid ichthyosaur[2]
Qianxisaurus[38]	Q. chajiangensis	A basal eosauropterygian, possibly related to pachypleurosaurs or nothosaurs
Tanystropheus	T. cf. hydroides[39]	A large tanystropheid, previously classified as T. cf. longobardicus[40]
	T. sp.	A small tanystropheid, previously classified as T. cf. longobardicus[40]
Wangosaurus[41]	W. brevirostris	A pistosauroid
Xinpusaurus	X. xingyiensis[42]	A thalattosauroid thalattosaur[29]
Yunguisaurus	Y. liae	A pistosauroid[43][24]