Errantia

Errantia is a diverse group of marine polychaete worms in the phylum Annelida. Traditionally a subclass of the paraphyletic class Polychaeta,^[4] it is currently regarded as a monophyletic group within the larger Pleistoannelida, composed of Errantia and Sedentaria.^[2] These worms are found worldwide in marine environments and brackish water.

Errantia Temporal range: Cambrian Stage 4–Recent PreꞒ Ꞓ O S D C P T J K Pg N [1]
Alitta virens
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Annelida
Clade:	Pleistoannelida
Subclass:	Errantia Audouin & H Milne Edwards 1832
Orders[2][3]
Aciculata Eunicida Phyllodocida Protodriliformia Polygordiidae Protodrilida

Phylogeny

The phylogeny of polychaetes is slowly being resolved. Errantia and Sedentaria are the two biggest clades of polychaetes, and together they compose clade Pleistoannelida.^[5] Two groups are nested within Errantia: Aciculata (Eunicida + Phyllodocida) and Protodriliformia (small meiofaunal worms such as the Protodrilida).^[6][3][2]

Historically, the order Amphinomida was part of this subclass. However, phylogenetic analyses place Amphinomida inside a basal clade with Sipunculida and Lobatocerebrum, and this clade is the sister group to Pleistoannelida.^[3]

	Amphinomida Lobatocerebrum Sipunculida
Pleistoannelida	Errantia Aciculata Eunicida Phyllodocida Protodriliformia Protodrilida Polygordiidae Sedentaria

Some taxa, such as Spintheridae and Myzostomida, are still difficult to place due to their long branching, but they likely belong to either Errantia or Sedentaria.^[3]

Classification

Historical

Errantia is, along with Sedentaria, one of the two old orders of the paraphyletic class "Polychaeta". In 1977 the zoologist Kristian Fauchald split Errantia into three orders: Phyllodocida, Amphinomida and Eunicida, giving way to this classification.^[7]

• Order Amphinomida – 2 families (Amphinomidae, Euphrosinidae)
• Order Eunicida
  • Superfamily Eunicea – 7 families (Onuphidae, Eunicidae, Lumbrineridae, Iphitimidae, Arabellidae, Lysaretidae, Dorvilleidae)
  • Superfamily incertae sedis – 2 families (Histriobdellidae, Ichthytomidae)
• Order Phyllodocida
  • Suborder Phyllodociformia – 4 families (Phyllodocidae, Alciopidae, Lopadorrhynchidae, Pontodoridae)
  • Suborder Aphroditiformia
    • Superfamily Aphroditacea – 6 families (Aphroditidae, Polynoidae, Polyodontidae, Pholoididae, Eulepethidae, Sigalionidae)
    • Superfamily Chrysopetalacea – 2 families (Chrysopetalidae, Palmyridae)
    • Superfamily Pisionacea – 1 family (Pisionidae)
  • Suborder Nereidiformia – 6 families (Hesionidae, Pilargiidae, Syllidae, Calamyzidae, Nereididae, Antonbmunidae)
  • Suborder Glyceriformia – 3 families (Glyceridae, Goniadidae, Lacydoniidae)
  • Suborder incertae sedis – 5 families (Iospilidae, Nephtyidae, Sphaerodoridae, Tomopteridae, Typhloscolecidae)