Clubiona

Clubiona is a genus of sac spiders that was first described by Pierre André Latreille in 1804.^[6]

Clubiona Temporal range: Palaeogene– Present PreꞒ Ꞓ O S D C P T J K Pg N
Clubiona trivialis
Male C. durbana
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Subphylum:	Chelicerata
Class:	Arachnida
Order:	Araneae
Infraorder:	Araneomorphae
Family:	Clubionidae
Genus:	Clubiona Latreille, 1804[1]
Type species
C. pallidula (Clerck, 1757)
Species
535, see text
Synonyms[1]
Anaclubiona Ono, 2010[2] Atalia Thorell, 1887[3] Bucliona Benoit, 1977[4] Hirtia Thorell, 1881[3] Tolophus Thorell, 1891[5]

Description

Species

As of October 2025^[update], this genus includes 535 species and five subspecies, found in Oceania, Africa, North America, the Caribbean, Asia, Europe, South America, Panama, and on Saint Helena.^[1]

• 
  female C. bevisi
• 
  male C. citricolor
• 
  C. huttoni
• 
  female C. pupillaris

These species have articles on Wikipedia:^[1]

• Clubiona abbajensis Strand, 1906 – Ethiopia, Somalia, Uganda, Rwanda, Zimbabwe, South Africa
• Clubiona abboti L. Koch, 1866 – Canada, United States
• Clubiona aducta Simon, 1932 – Portugal, Spain, Italy
• Clubiona africana Lessert, 1921 – Tanzania, South Africa
• Clubiona alpicola Kulczyński, 1882 – France, Central Europe, Italy, Romania, Bulgaria, Ukraine
• Clubiona andreinii Caporiacco, 1936 – Italy
• Clubiona annuligera Lessert, 1929 – DR Congo, Malawi, Mozambique, South Africa
• Clubiona aspidiphora Simon, 1910 – Namibia, South Africa
• Clubiona bevisi Lessert, 1923 – South Africa
• Clubiona biaculeata Simon, 1897 – South Africa
• Clubiona bishopi Edwards, 1958 – Canada, United States
• Clubiona blesti Forster, 1979 – New Zealand
• Clubiona brevipes Blackwall, 1841 – Europe, Turkey, Caucasus, Japan?
• Clubiona cada Forster, 1979 – New Zealand
• Clubiona cambridgei L. Koch, 1873 – New Zealand
• Clubiona canadensis Emerton, 1890 – Canada, United States
• Clubiona capensis Simon, 1897 – South Africa
• Clubiona chathamensis Simon, 1905 – New Zealand (Chatham Is.)
• Clubiona citricolor Lawrence, 1952 – South Africa
• Clubiona clima Forster, 1979 – New Zealand
• Clubiona comta C. L. Koch, 1839 – Europe, North Africa, Turkey, Caucasus
• Clubiona consensa Forster, 1979 – New Zealand
• Clubiona contrita Forster, 1979 – New Zealand
• Clubiona convoluta Forster, 1979 – New Zealand
• Clubiona corticalis (Walckenaer, 1802) – Europe, Turkey, Caucasus
• Clubiona delicata Forster, 1979 – New Zealand
• Clubiona deterrima Strand, 1904 – Norway
• Clubiona durbana Roewer, 1951 – South Africa
• Clubiona facilis O. Pickard-Cambridge, 1911 – Britain (presumably imported)
• Clubiona frutetorum L. Koch, 1867 – Europe to Central Asia
• Clubiona germanica Thorell, 1871 – Europe, Caucasus, Russia (Europe to Far East), Central Asia
• Clubiona godfreyi Lessert, 1921 – Uganda, South Africa
• Clubiona helva Simon, 1897 – South Africa
• Clubiona hilaris Simon, 1878 – Mountains of Spain, France, Italy, Austria, Switzerland, North Macedonia and Romania
• Clubiona huttoni Forster, 1979 – New Zealand
• Clubiona hwanghakensis Paik, 1990 – Korea
• Clubiona johnsoni Gertsch, 1941 – Canada, United States
• Clubiona kastoni Gertsch, 1941 – Alaska, Canada, United States
• Clubiona kiboschensis Lessert, 1921 – Tanzania, South Africa
• Clubiona kulczynskii Lessert, 1905 – North America, Europe, Kazakhstan, Russia (Europe to Far East), Japan
• Clubiona lawrencei Roewer, 1951 – Zimbabwe, South Africa
• Clubiona limpida Simon, 1897 – South Africa
• Clubiona lutescens Westring, 1851 – Europe, Turkey, Caucasus, Russia (Europe to Far East), Iran, Kazakhstan, Korea, Japan. Introduced to North America
• Clubiona maritima L. Koch, 1867 – Canada, United States, Caribbean
• Clubiona marmorata L. Koch, 1866 – France to Ukraine and Turkey
• Clubiona marna Roddy, 1966 – United States
• Clubiona mimula Chamberlin, 1928 – Canada, United States
• Clubiona moesta Banks, 1896 – China, Alaska, Canada, United States
• Clubiona mykolai Mikhailov, 2003 – Bulgaria, Ukraine
• Clubiona natalica Simon, 1897 – South Africa
• Clubiona nemorum Ledoux, 2004 – Réunion
• Clubiona nigromaculosa Blackwall, 1877 – Seychelles, Réunion
• Clubiona nollothensis Simon, 1910 – South Africa
• Clubiona norvegica Strand, 1900 – North America, Europe, Russia (Europe to West Siberia)
• Clubiona obesa Hentz, 1847 – Canada, United States
• Clubiona pacifica Banks, 1896 – Alaska, Canada, United States
• Clubiona pallidula (Clerck, 1757) – Europe, Turkey, Caucasus, Russia (Europe to Far East), Kazakhstan, Central Asia. Introduced to North America (type species)
• Clubiona peculiaris L. Koch, 1873 – New Zealand
• Clubiona pikei Gertsch, 1941 – Canada, United States
• Clubiona pongolensis Lawrence, 1952 – South Africa
• Clubiona producta Forster, 1979 – New Zealand
• Clubiona pseudoneglecta Wunderlich, 1994 – Morocco, Algeria, Europe, Turkey, Caucasus, Iran
• Clubiona pupillaris Lawrence, 1938 – South Africa
• Clubiona pygmaea Banks, 1892 – Canada, United States
• Clubiona revillioidi Lessert, 1936 – Mozambique, South Africa, Eswatini
• Clubiona riparia L. Koch, 1866 – Russia (Urals to Far East), Mongolia, China, Japan, North America
• Clubiona robusta L. Koch, 1873 – Australia
• Clubiona rosserae Locket, 1953 – Britain, France, Netherlands, Poland, Slovakia, Hungary, Romania, Serbia
• Clubiona rumpiana Lawrence, 1952 – South Africa
• Clubiona saltuum Kulczyński, 1898 – Austria
• Clubiona saxatilis L. Koch, 1867 – France to Ukraine and south-eastern Europe
• Clubiona scatula Forster, 1979 – New Zealand
• Clubiona sigillata Lawrence, 1952 – South Africa
• Clubiona silvestris Deeleman-Reinhold, 2001 – Malaysia (Borneo)
• Clubiona sparassella Strand, 1909 – South Africa
• Clubiona spiralis Emerton, 1909 – Canada, United States
• Clubiona subsultans Thorell, 1875 – Europe, Russia (Europe to South Siberia), Japan
• Clubiona subtilis L. Koch, 1867 – Europe, Russia (Europe to Far East), Kyrgyzstan, Korea
• Clubiona subtrivialis Strand, 1906 – Ethiopia, East Africa, South Africa
• Clubiona terrestris Westring, 1851 – Azores, Europe, Turkey
• Clubiona torta Forster, 1979 – New Zealand
• Clubiona trivialis C. L. Koch, 1843 – North America, Europe, Russia (Europe to Far East), China, Japan
• Clubiona umbilensis Lessert, 1923 – South Africa
• Clubiona vachoni Lawrence, 1952 – South Africa
• Clubiona valens Simon, 1897 – South Africa

Complete species list as of October 2025[update]

C. abbajensis Strand, 1906 – Ethiopia, Somalia, Uganda, Rwanda, Zimbabwe, South Africa C. a. kibonotensis Lessert, 1921 – DR Congo, Uganda, Tanzania C. abboti L. Koch, 1866 – Canada, United States C. a. abbotoides Chamberlin & Ivie, 1946 – United States C. aberrans Dankittipakul, 2012 – Thailand C. abnormis Dankittipakul, 2008 – China, Thailand, Laos C. acanthocnemis Simon, 1906 – India C. achilles Hogg, 1896 – Australia (Central) C. acies Nicolet, 1849 – Chile C. aciformis Zhang & Hu, 1991 – China C. aculeata Zhang, Zhu & Song, 2007 – China C. adjacens Gertsch & Davis, 1936 – United States C. aducta Simon, 1932 – Portugal, Spain, Italy C. africana Lessert, 1921 – Tanzania, South Africa C. akagiensis Hayashi, 1985 – Japan C. alexeevi Mikhailov, 1990 – Russia (Caucasus) C. aliceae Chickering, 1937 – Panama C. allotorta Dankittipakul & Singtripop, 2008 – Thailand C. alluaudi Simon, 1898 – Mauritius C. alpicola Kulczyński, 1882 – France, Central Europe, Italy, Romania, Bulgaria, Ukraine C. alticola Dankittipakul & Singtripop, 2008 – Thailand C. altissimoides Liu, Yan, Griswold & Ubick, 2007 – China C. altissimus Hu, 2001 – China C. alveolata L. Koch, 1873 – Samoa, Funafuti, Marquesas Is. Hawaii C. amurensis Mikhailov, 1990 – Russia (Far East), Japan C. analis Thorell, 1895 – India, Bangladesh, Myanmar C. andreinii Caporiacco, 1936 – Italy C. angulata Dondale & Redner, 1976 – Canada C. annuligera Lessert, 1929 – DR Congo, Malawi, Mozambique, South Africa C. ansa Jang, Bae, Yoo, Lee & Kim, 2021 – Korea C. anwarae Biswas & Raychaudhuri, 1996 – Bangladesh C. apiata Urquhart, 1893 – Australia (Tasmania) C. apiculata Dankittipakul & Singtripop, 2014 – Malaysia (Borneo) C. applanata Liu, Yan, Griswold & Ubick, 2007 – China C. aspidiphora Simon, 1910 – Namibia, South Africa C. asrevida Ono, 1992 – Taiwan C. auberginosa Zhang, Yin, Bao & Kim, 1997 – China C. australiaca Kolosváry, 1934 – Australia (New South Wales) C. bachmaensis Ono, 2009 – Vietnam C. bagerhatensis Biswas & Raychaudhuri, 1996 – Bangladesh C. baimaensis Song & Zhu, 1991 – China C. baishishan Zhang, Zhu & Song, 2003 – China C. bakurovi Mikhailov, 1990 – Russia (Far East), China, Korea C. bandoi Hayashi, 1995 – Japan C. banna Yu & Li, 2021 – China C. basarukini Mikhailov, 1990 – Russia (South Siberia, Far East), Mongolia, Japan C. bashkirica Mikhailov, 1992 – Russia (Urals, West Siberia) C. bengalensis Biswas, 1984 – India C. bevisi Lessert, 1923 – South Africa C. bi Zhang, Zhong & Gong, 2024 – China C. biaculeata Simon, 1897 – South Africa C. bicornis Yu & Li, 2019 – China C. bicuspidata Wu & Zhang, 2014 – China C. bidactylina Wu, Chen & Zhang, 2023 – China C. biembolata Deeleman-Reinhold, 2001 – Malaysia (Borneo), Indonesia (Sumatra) C. bifissurata Kritscher, 1966 – New Caledonia C. biforamina Liu, Peng & Yan, 2016 – China C. bifurcata Zhang, Yu & Zhong, 2018 – China C. bilobata Dhali, Roy, Saha & Raychaudhuri, 2016 – India C. bipinnata Yu, Zhang & Chen, 2017 – China C. bishopi Edwards, 1958 – Canada, United States C. blesti Forster, 1979 – New Zealand C. bomiensis Zhang & Zhu, 2009 – China C. boxaensis B. Biswas & K. Biswas, 1992 – India C. brevipes Blackwall, 1841 – Europe, Turkey, Caucasus, Japan? C. brevispina Huang & Chen, 2012 – Taiwan C. bryantae Gertsch, 1941 – Alaska, Canada, United States C. bucera Yang, Ma & Zhang, 2011 – China C. bukaea (Barrion & Litsinger, 1995) – Philippines C. cada Forster, 1979 – New Zealand C. caerulescens L. Koch, 1867 – Europe, Russia (Europe to Far East), Kazakhstan, China, Japan C. californica Fox, 1938 – United States C. calycina Wu & Zhang, 2014 – China C. cambridgei L. Koch, 1873 – New Zealand C. camela Wu, Chen & Zhang, 2023 – China C. campylacantha Dankittipakul, 2008 – Thailand C. canaca Berland, 1930 – New Caledonia C. canadensis Emerton, 1890 – Canada, United States C. canberrana Dondale, 1966 – Australia (New South Wales) C. candefacta Nicolet, 1849 – Chile C. caohai Zhang & Yu, 2020 – China C. capensis Simon, 1897 – South Africa C. catawba Gertsch, 1941 – United States C. caucasica Mikhailov & Otto, 2017 – Turkey, Caucasus (Russia, Georgia, Armenia, Azerbaijan) C. chabarovi Mikhailov, 1991 – Russia (Far East) C. chakrabartei Majumder & Tikader, 1991 – India C. charitonovi Mikhailov, 1990 – Russia (South Siberia, Far East) C. charleneae Barrion & Litsinger, 1995 – Philippines C. chathamensis Simon, 1905 – New Zealand (Chatham Is.) C. cheni Yu & Li, 2019 – China C. chevalieri Berland, 1936 – Cape Verde C. chikunii Hayashi, 1986 – Japan C. chippewa Gertsch, 1941 – Canada, United States C. circulata Zhang & Yin, 1998 – China, Laos, Thailand C. cirrosa Ono, 1989 – Japan (Ryukyu Is.) C. citricolor Lawrence, 1952 – South Africa C. clima Forster, 1979 – New Zealand C. cochlearis Yu & Li, 2019 – China C. cochleata Wang, Wu & Zhang, 2015 – China C. complicata Banks, 1898 – Mexico C. comta C. L. Koch, 1839 – Europe, North Africa, Turkey, Caucasus C. concinna (Thorell, 1887) – Myanmar C. congentilis Kulczyński, 1913 – Slovakia, Ukraine, Russia (Europe to South Siberia), Kazakhstan C. conica Dankittipakul & Singtripop, 2014 – Malaysia (Borneo) C. consensa Forster, 1979 – New Zealand C. contaminata O. Pickard-Cambridge, 1872 – Israel C. contrita Forster, 1979 – New Zealand C. convoluta Forster, 1979 – New Zealand C. cordata Zhang & Zhu, 2009 – China C. coreana Paik, 1990 – Russia (Far East), China, Korea, Japan C. corrugata Bösenberg & Strand, 1906 – Russia (Far East), China, Taiwan, Korea, Japan, Thailand C. corticalis (Walckenaer, 1802) – Europe, Turkey, Caucasus C. c. concolor Kulczyński, 1897 – Hungary C. c. nigra Simon, 1878 – France C. cultrata Dankittipakul & Singtripop, 2014 – Indonesia (Borneo) C. cycladata Simon, 1909 – Australia (Western Australia) C. cylindrata Liu, Yan, Griswold & Ubick, 2007 – China C. cylindriformis Dankittipakul & Singtripop, 2014 – Malaysia (Borneo) C. dactylina Liu, Peng & Yan, 2016 – China C. dakong Zhang & Yu, 2020 – China C. damirkovaci Deeleman-Reinhold, 2001 – Malaysia (peninsula) C. debilis Nicolet, 1849 – Chile C. deletrix O. Pickard-Cambridge, 1885 – Pakistan, India, China?, Taiwan? Japan? C. delicata Forster, 1979 – New Zealand C. dengpao Yu & Li, 2021 – China C. denticulata Dhali, Roy, Saha & Raychaudhuri, 2016 – India C. desecheonis Petrunkevitch, 1930 – Puerto Rico C. deterrima Strand, 1904 – Norway C. dichotoma Wang, Chen & Z. S. Zhang, 2018 – China C. didentata Zhang & Yin, 1998 – China C. digitata Dankittipakul, 2012 – Thailand C. dikita Barrion & Litsinger, 1995 – Philippines C. diversa O. Pickard-Cambridge, 1862 – Europe, Turkey, Caucasus, Russia (Europe to Far East), Kazakhstan, Pakistan, Korea, Japan C. dorni Sarkar, Quasin & Siliwal, 2023 – India (Himalayas) C. drassodes O. Pickard-Cambridge, 1874 – India, Bangladesh, China C. dunini Mikhailov, 2003 – Russia (Far East) C. duoconcava Zhang & Hu, 1991 – China C. durbana Roewer, 1951 – South Africa C. dyasia Gertsch, 1941 – United States C. dysderiformis (Guérin, 1838) – Indonesia (New Guinea) C. elaphines Urquhart, 1893 – Australia (Tasmania) C. ericius Chrysanthus, 1967 – Indonesia (New Guinea) C. eskovi Mikhailov, 1995 – Russia (Far East) C. estes Edwards, 1958 – United States C. esuriens Thorell, 1897 – Myanmar C. evoronensis Mikhailov, 1995 – Russia (Far East) C. excavata (Rainbow, 1920) – Australia (Lord Howe Is.) C. excisa O. Pickard-Cambridge, 1898 – Mexico C. ezoensis Hayashi, 1987 – Russia (Far East), Japan C. facilis O. Pickard-Cambridge, 1911 – Britain (presumably imported) C. falcata Tang, Song & Zhu, 2005 – China, Mongolia C. falciforma Liu, Peng & Yan, 2016 – China C. fanjingshan Wang, Chen & Z. S. Zhang, 2018 – China C. femorocalcarata Huang & Chen, 2012 – Taiwan C. filicata O. Pickard-Cambridge, 1874 – Pakistan, India, Bangladesh, Myanmar, Thailand, Laos, China, Taiwan C. filifera Dankittipakul, 2008 – Thailand C. filoramula Zhang & Yin, 1998 – China C. flammaformis L. F. Li, Liu, B. Li & Peng, 2023 – China C. flavocincta Nicolet, 1849 – Chile C. forcipa Yang, Song & Zhu, 2003 – China C. frisia Wunderlich & Schuett, 1995 – Europe to Central Asia C. frutetorum L. Koch, 1867 – Europe to Central Asia C. furcata Emerton, 1919 – Russia (Middle Siberia to Far East), Alaska, Canada, United States, Mexico C. fusoidea Zhang, 1992 – China C. fuzhouensis Gong, 1985 – China C. gallagheri Barrion & Litsinger, 1995 – Indonesia (Java) C. germanica Thorell, 1871 – Europe, Caucasus, Russia (Europe to Far East), Central Asia C. gertschi Edwards, 1958 – United States C. gilva O. Pickard-Cambridge, 1872 – Israel C. giulianetti Rainbow, 1898 – Papua New Guinea C. glatiosa Saito, 1934 – Japan C. globosa Wang, Chen & Z. S. Zhang, 2018 – China C. godfreyi Lessert, 1921 – Uganda, South Africa C. golovatchi Mikhailov, 1990 – Russia (Europe), Caucasus, Turkey, Iran C. gongi Zhang, Yin, Bao & Kim, 1997 – China C. gongshan He, Liu & Zhang, 2016 – China C. grucollaris Yu, Zhang & Chen, 2017 – China C. guianensis Caporiacco, 1947 – Guyana C. haeinsensis Paik, 1990 – Russia (Far East), China, Korea, Japan C. haplotarsus Simon, 1909 – São Tomé and Príncipe C. hatamensis (Thorell, 1881) – Indonesia (New Guinea) C. haupti Tang, Song & Zhu, 2005 – China C. hedini Schenkel, 1936 – China C. helenae Mikhailov, 2003 – Russia (Far East) C. helva Simon, 1897 – South Africa C. heteroducta Zhang & Yin, 1998 – China C. heterosaca Yin, Yan, Gong & Kim, 1996 – China C. hexadentata Dhali, Roy, Saha & Raychaudhuri, 2016 – India C. hilaris Simon, 1878 – Mountains of Spain, France, Italy, Austria, Switzerland, North Macedonia and Romania C. hindu Deeleman-Reinhold, 2001 – Indonesia (Bali) C. hitchinsi Saaristo, 2002 – St. Helena, Ascension Island, Seychelles, French Polynesia (Tuamotu) C. hoffmanni Schenkel, 1937 – Madagascar C. hooda Dong & Zhang, 2016 – China C. huaban Xin, Zhang, Li, Zheng & Yu, 2020 – China C. hugispaa Barrion & Litsinger, 1995 – Philippines C. hugisva Barrion & Litsinger, 1995 – Philippines C. huiming Wang, F. Zhang & Z. S. Zhang, 2018 – China C. hummeli Schenkel, 1936 – China C. hundeshageni Strand, 1907 – Indonesia (Moluccas) C. huntianling Yu & Li, 2025 – China C. huojianqiang Yu & Li, 2025 – China C. huttoni Forster, 1979 – New Zealand C. hwanghakensis Paik, 1990 – Korea C. hyrcanica Mikhailov, 1990 – Azerbaijan, Iran C. hysgina Simon, 1889 – India C. hystrix Berland, 1938 – Indonesia (Lesser Sunda Is.), Vanuatu C. iharai Ono, 1995 – Japan C. ikedai Ono, 1992 – Japan C. inaensis Hayashi, 1989 – Japan C. inquilina Deeleman-Reinhold, 2001 – Malaysia (Borneo) C. insulana Ono, 1989 – Taiwan, Japan (Ryukyu Is.) C. interjecta L. Koch, 1879 – Russia (West Siberia to Far East), Mongolia, China C. irinae Mikhailov, 1991 – Russia (Far East), China, Korea C. jaegeri Ono, 2011 – Palau Is. C. janae Edwards, 1958 – United States C. japonica L. Koch, 1878 – Russia (Sakhalin, Kurile Is.), China, Korea, Taiwan, Japan C. japonicola Bösenberg & Strand, 1906 – Russia (Far East), Korea, Japan, China, Taiwan, Philippines, Indonesia C. jiandan Yu & Li, 2019 – China C. jiugong Yu & Zhong, 2021 – China C. jiulongensis Zhang, Yin & Kim, 1996 – China C. johnsoni Gertsch, 1941 – Canada, United States C. juvenis Simon, 1878 – Europe, Iran, Uzbekistan C. kagani Gertsch, 1941 – United States C. kai Jäger & Dankittipakul, 2010 – China, Laos C. kaltenbachi Kritscher, 1966 – New Caledonia C. kapataganensis Barrion & Litsinger, 1995 – Philippines C. kasanensis Paik, 1990 – Korea, Japan C. kastoni Gertsch, 1941 – Alaska, Canada, United States C. kasurensis Mukhtar & Mushtaq, 2005 – Pakistan C. katioryza Barrion & Litsinger, 1995 – Philippines C. kawitpaaia (Barrion & Litsinger, 1995) – Philippines C. kayashimai Ono, 1994 – Taiwan C. kiboschensis Lessert, 1921 – Tanzania, South Africa C. kigabensis Strand, 1915 – East Africa C. kimyongkii Paik, 1990 – Russia (Far East), China, Korea C. kiowa Gertsch, 1941 – Canada, United States, Mexico C. komissarovi Mikhailov, 1992 – Russia (Far East), Korea C. kowong Chrysanthus, 1967 – Indonesia (New Guinea) C. krisisensis Barrion & Litsinger, 1995 – Philippines, Indonesia (Borneo) C. kropfi Zhang, Zhu & Song, 2003 – China C. kuanshanensis Ono, 1994 – Taiwan C. kularensis Marusik & Koponen, 2002 – Russia (north-eastern Siberia, Far East) C. kulczynskii Lessert, 1905 – North America, Europe, Kazakhstan, Russia (Europe to Far East), Japan C. kumadaorum Ono, 1992 – Japan C. kunashirensis Mikhailov, 1990 – Russia (Sakhalin, Kurile Is.), Japan C. kurenshikovi Mikhailov, 1995 – Russia (Far East) C. kurilensis Bösenberg & Strand, 1906 – Russia (Far East), Korea, Japan, China, Taiwan C. kurosawai Ono, 1986 – Korea, Japan, China, Taiwan C. kuu Jäger & Dankittipakul, 2010 – Laos C. lala Jäger & Dankittipakul, 2010 – China, Laos C. lamellaris Zhang, Yu & Zhong, 2018 – China C. lamina Zhang, Zhu & Song, 2007 – China C. langei Mikhailov, 1991 – Russia (Far East) C. latericia Kulczyński, 1926 – Russia (Middle Siberia to Far East), Mongolia, Alaska C. laticeps O. Pickard-Cambridge, 1885 – Pakistan C. latitans Pavesi, 1883 – Ethiopia, Somalia, Kenya C. laudabilis Simon, 1909 – Australia (Western Australia) C. lawrencei Roewer, 1951 – Zimbabwe, South Africa C. lena Bösenberg & Strand, 1906 – China, Korea, Japan C. leonilae Barrion & Litsinger, 1995 – Philippines C. leptosa Zhang, Yin, Bao & Kim, 1997 – China C. liachviana Mcheidze, 1997 – Caucasus (Russia, Georgia, Armenia, Azerbaijan), Iran, Turkmenistan C. liminal Sherwood, 2025 – China, Korea C. limpida Simon, 1897 – South Africa C. linea Xie, Yin, Yan & Kim, 1996 – China C. linzhiensis Hu, 2001 – China C. lirata Yang, Song & Zhu, 2003 – China C. littoralis Banks, 1895 – Canada, United States C. logunovi Mikhailov, 1990 – Russia (Far East) C. longipes Nicolet, 1849 – Chile C. longyangensis Guo, Li & Zhang, 2025 – China C. luapalana Giltay, 1935 – DR Congo C. lucida He, Liu & Zhang, 2016 – China C. ludhianaensis Tikader, 1976 – India, Bangladesh C. lui X. Y. Zhang, Li & Z. S. Zhang, 2025 – China C. lutescens Westring, 1851 – Europe, Turkey, Caucasus, Russia (Europe to Far East), Iran, Kazakhstan, Korea, Japan. Introduced to North America C. lyriformis Song & Zhu, 1991 – China C. maculata Roewer, 1951 – Australia (Queensland) C. mahensis Simon, 1893 – Seychelles C. maipai Jäger & Dankittipakul, 2010 – Thailand C. mandschurica Schenkel, 1953 – Russia (Far East), China, Korea, Japan C. manshanensis Zhu & An, 1988 – China C. maracandica Kroneberg, 1875 – Uzbekistan C. maritima L. Koch, 1867 – Canada, United States, Caribbean C. marmorata L. Koch, 1866 – France to Ukraine and Turkey C. marna Roddy, 1966 – United States C. marusiki Mikhailov, 1990 – Russia (Far East) C. maya Hayashi & Yoshida, 1991 – Japan C. maysangarta Barrion & Litsinger, 1995 – Philippines C. mayumiae Ono, 1993 – Russia (Far East), Korea, Japan C. mazandaranica Mikhailov, 2003 – Azerbaijan, Iran C. medog Zhang, Zhu & Song, 2007 – China C. melanosticta Thorell, 1890 – India, China, Myanmar, Thailand, Laos, Thailand, Indonesia, Papua New Guinea C. menglun Yu & Li, 2021 – China C. meraukensis Chrysanthus, 1967 – Malaysia, Indonesia (New Guinea) C. microsapporensis Mikhailov, 1990 – Russia (Far East), Korea C. mii Yu & Li, 2021 – China C. mikhailovi Deeleman-Reinhold, 2001 – Indonesia (Java) C. milingae Barrion-Dupo, Barrion & Heong, 2013 – China (Hainan) C. mimula Chamberlin, 1928 – Canada, United States C. minima (Ono, 2010) – Korea, Japan C. minuscula Nicolet, 1849 – Chile C. minuta Nicolet, 1849 – Chile C. mixta Emerton, 1890 – Canada, United States C. modesta L. Koch, 1873 – Australia (Queensland) C. moesta Banks, 1896 – China, Alaska, Canada, United States C. moralis Song & Zhu, 1991 – China, Taiwan C. mordica O. Pickard-Cambridge, 1898 – Mexico C. mujibari Biswas & Raychaudhuri, 1996 – Bangladesh C. multidentata Liu, Peng & Yan, 2016 – China C. multiprocessa Guo, Li & Zhang, 2025 – China C. munda Thorell, 1887 – Myanmar C. munis Simon, 1909 – Australia (Western Australia) C. mutata Gertsch, 1941 – Canada, United States C. mutilata Bösenberg & Strand, 1906 – Japan C. mykolai Mikhailov, 2003 – Bulgaria, Ukraine C. nataliae Trilikauskas, 2007 – Russia (Far East) C. natalica Simon, 1897 – South Africa C. neglecta O. Pickard-Cambridge, 1862 – Europe, Turkey, Caucasus, Russia (Europe to South Siberia), Iran, Central Asia, China, Korea C. neglectoides Bösenberg & Strand, 1906 – China, Korea, Japan C. nemorum Ledoux, 2004 – Réunion C. nenilini Mikhailov, 1995 – Russia (South Siberia) C. neocaledonica Berland, 1924 – New Caledonia C. newnani Ivie & Barrows, 1935 – United States C. nezha Yu & Li, 2025 – China C. nicholsi Gertsch, 1941 – Canada, United States C. nicobarensis Tikader, 1977 – India (Nicobar Is.) C. nigromaculosa Blackwall, 1877 – Seychelles, Réunion C. nilgherina Simon, 1906 – India C. ningpoensis Schenkel, 1944 – China C. nollothensis Simon, 1910 – South Africa C. norvegica Strand, 1900 – North America, Europe, Russia (Europe to West Siberia) C. notabilis L. Koch, 1873 – Australia (Queensland) C. obesa Hentz, 1847 – Canada, United States C. oceanica Ono, 2011 – Japan C. octoginta Dankittipakul, 2008 – Thailand C. odelli Edwards, 1958 – United States C. odesanensis Paik, 1990 – Russia (Far East), China, Korea C. ogatai Ono, 1995 – Japan C. oligerae Mikhailov, 1995 – Russia (Far East) C. opeongo Edwards, 1958 – Canada C. orientalis Mikhailov, 1995 – North Korea C. oteroana Gertsch, 1941 – United States C. ovalis Zhang, 1991 – China C. pacifica Banks, 1896 – Alaska, Canada, United States C. paenuliformis (Strand, 1916) – Ghana C. pahilistapyasea Barrion & Litsinger, 1995 – Thailand, Indonesia (Borneo), Philippines C. paiki Mikhailov, 1991 – Russia (Far East) C. pala Deeleman-Reinhold, 2001 – Indonesia (Moluccas) C. pallidula (Clerck, 1757) – Europe, Turkey, Caucasus, Russia (Europe to Far East), Kazakhstan, Central Asia. Introduced to North America (type species) C. pandalira (Barrion-Dupo, Barrion & Heong, 2013) – China C. pantherina Chrysanthus, 1967 – Indonesia (New Guinea) C. papillata Schenkel, 1936 – Russia (Far East), China, Korea C. papuana Chrysanthus, 1967 – Indonesia (New Guinea) C. paralena Mikhailov, 1995 – North Korea C. parallela Hu & Li, 1987 – China C. paranghinlalakirta Barrion & Litsinger, 1995 – Philippines C. parangunikarta Barrion & Litsinger, 1995 – Philippines C. parconcinna Deeleman-Reinhold, 2001 – China, Thailand, Indonesia (Borneo) C. parva Seo, 2018 – Korea C. parvula Saito, 1933 – Japan C. peculiaris L. Koch, 1873 – New Zealand C. phansa Strand, 1911 – Indonesia (Aru Is.) C. phragmitis C. L. Koch, 1843 – Morocco, Algeria, Europe, Caucasus, Russia (Europe to Far East), Iran, Central Asia, China, Korea C. phragmitoides Schenkel, 1963 – China C. pianmaensis Wang, Wu & Zhang, 2015 – China C. picturata Deeleman-Reinhold, 2001 – Indonesia (Bali) C. pikei Gertsch, 1941 – Canada, United States C. pila Dhali, Roy, Saha & Raychaudhuri, 2016 – India C. plumbi Gertsch, 1941 – United States C. pogonias Simon, 1906 – India C. pollicaris Wu, Zheng & Zhang, 2015 – China C. pomoa Gertsch, 1941 – United States C. pongolensis Lawrence, 1952 – South Africa C. pototanensis Barrion & Litsinger, 1995 – Philippines C. praematura Emerton, 1909 – Russia (Far East), Alaska, Canada, United States C. procera Chrysanthus, 1967 – Indonesia (New Guinea) C. procteri Gertsch, 1941 – United States C. producta Forster, 1979 – New Zealand C. propinqua L. Koch, 1879 – Russia (Middle Siberia to Far East), North Korea, China C. proszynskii Mikhailov, 1995 – North Korea C. pruvotae Berland, 1930 – New Caledonia C. pseudocordata Dhali, Roy, Saha & Raychaudhuri, 2016 – India C. pseudogermanica Schenkel, 1936 – Russia (Far East), China, Korea, Japan C. pseudomaxillata Hogg, 1915 – Indonesia (New Guinea) C. pseudoneglecta Wunderlich, 1994 – Morocco, Algeria, Europe, Turkey, Caucasus, Iran C. pseudopteroneta Raven & Stumkat, 2002 – Australia (Queensland) C. pseudosaxatilis Mikhailov, 1992 – Russia (Central Asia, South Siberia), Kazakhstan C. pseudosimilis Mikhailov, 1990 – Algeria, Portugal, Greece (Crete), Turkey, Caucasus C. pterogona Yang, Song & Zhu, 2003 – China C. puera Nicolet, 1849 – Chile C. pupillaris Lawrence, 1938 – South Africa C. pupula Thorell, 1897 – Myanmar C. pygmaea Banks, 1892 – Canada, United States C. pyrifera Schenkel, 1936 – China C. qiankunquan Yu & Li, 2025 – China C. qianlei J. S. Zhang, F. Zhang & Yu, 2022 – China C. qini Tang, Song & Zhu, 2005 – China C. qiyunensis Xu, Yang & Song, 2003 – China C. quebecana Dondale & Redner, 1976 – Canada, United States C. rainbowi Roewer, 1951 – Australia (Lord Howe Is.) C. rama Dankittipakul & Singtripop, 2008 – India, Thailand, China C. ramoiensis (Thorell, 1881) – Papua New Guinea C. rava Simon, 1886 – Senegal C. reclusa O. Pickard-Cambridge, 1863 – Europe, Turkey, Russia (Europe to South Siberia), Georgia, Kazakhstan C. reichlini Schenkel, 1944 – China, Taiwan? Japan? C. revillioidi Lessert, 1936 – Mozambique, South Africa, Eswatini C. rhododendri Barrows, 1945 – United States C. rileyi Gertsch, 1941 – United States C. riparia L. Koch, 1866 – Russia (Urals to Far East), Mongolia, China, Japan, North America C. risbeci Berland, 1930 – New Caledonia C. rivalis Pavesi, 1883 – Ethiopia C. robusta L. Koch, 1873 – Australia C. roeweri Caporiacco, 1940 – Ethiopia C. rosserae Locket, 1953 – Britain, France, Netherlands, Poland, Slovakia, Hungary, Romania, Serbia C. rostrata Paik, 1985 – Russia (Far East), China, Korea, Japan C. rothschildi Berland, 1922 – Ethiopia C. rouqiu Yu & Li, 2025 – China C. rumpiana Lawrence, 1952 – South Africa C. rybini Mikhailov, 1992 – Kazakhstan, Kyrgyzstan C. ryukyuensis Ono, 1989 – Japan (Ryukyu Is.) C. saltitans Emerton, 1919 – Canada, United States C. saltuum Kulczyński, 1898 – Austria C. samoensis Berland, 1929 – Samoa, French Polynesia (Society Is., Austral Is.: Rapa) C. sapporensis Hayashi, 1986 – Russia (Far East), Korea, Japan C. saurica Mikhailov, 1992 – Kazakhstan C. savesi Berland, 1930 – New Caledonia C. saxatilis L. Koch, 1867 – France to Ukraine and south-eastern Europe C. scandens Deeleman-Reinhold, 2001 – Malaysia (Borneo) C. scatula Forster, 1979 – New Zealand C. scenica Nicolet, 1849 – Chile C. semicircularis Tang, Song & Zhu, 2005 – China, Korea C. sertungensis Hayashi, 1996 – Indonesia (Krakatau) C. shillongensis Majumder & Tikader, 1991 – India C. shuangsi Yu & Li, 2021 – China C. sichotanica Mikhailov, 2003 – Russia (Far East) C. sigillata Lawrence, 1952 – South Africa C. silvestris Deeleman-Reinhold, 2001 – Malaysia (Borneo) C. similis L. Koch, 1867 – Europe, Turkey, Caucasus C. sjostedti Lessert, 1921 – Tanzania C. s. spinigera Lessert, 1921 – Tanzania C. sopaikensis Paik, 1990 – Russia (Far East), Korea C. sparassella Strand, 1909 – South Africa C. spiralis Emerton, 1909 – Canada, United States C. stagnatilis Kulczyński, 1897 – Europe, Caucasus, Russia (Europe to South Siberia), Central Asia C. stiligera Deeleman-Reinhold, 2001 – Indonesia (Sumatra) C. straminea O. Pickard-Cambridge, 1872 – Israel C. subapplanata Wang, Chen & Z. S. Zhang, 2018 – China C. subasrevida Yu & Li, 2019 – China C. subborealis Mikhailov, 1992 – Russia (South Siberia, Far East), Mongolia C. subcylindrata Wang, Chen & Z. S. Zhang, 2018 – China C. subdidentata Yu & Li, 2021 – China C. subhuiming Wu, Chen & Zhang, 2023 – China C. subkuu Yu & Li, 2019 – China C. submaculata (Thorell, 1891) – India (Nicobar Is.) C. submoralis Wu, Zheng & Zhang, 2015 – China C. subnotabilis Strand, 1907 – Australia C. subparallela Zhang, Zhu & Song, 2007 – China C. subquebecana Yu & Li, 2019 – China C. subrama Yu & Li, 2019 – China C. subrostrata Zhang & Hu, 1991 – China C. subsultans Thorell, 1875 – Europe, Russia (Europe to South Siberia), Japan C. subtilis L. Koch, 1867 – Europe, Russia (Europe to Far East), Kyrgyzstan, Korea C. subtongi Yu & Li, 2021 – China C. subtrivialis Strand, 1906 – Ethiopia, East Africa, South Africa C. subyaginumai Yu & Li, 2019 – China C. subyangmingensis Gan & Wang, 2020 – China C. suthepica Dankittipakul, 2008 – China, Thailand C. tabupumensis Petrunkevitch, 1914 – Myanmar C. taiwanica Ono, 1994 – China, Taiwan C. tangi Liu, Peng & Yan, 2016 – China C. tanikawai Ono, 1989 – China, Taiwan, Japan (Ryukyu Is.) C. tateyamensis Hayashi, 1989 – Japan C. tenera (Thorell, 1890) – Indonesia (Sumatra, Java) C. tengchong Zhang, Zhu & Song, 2007 – China C. ternatensis (Thorell, 1881) – Indonesia (Moluccas) C. terrestris Westring, 1851 – Azores, Europe, Turkey C. theoblicki Yu & Li, 2019 – China C. thorelli Roewer, 1951 – Indonesia (Sumatra) C. tiane Yu & Li, 2019 – China C. tianpingshan L. F. Li, Liu, B. Li & Peng, 2023 – China C. tiantongensis Zhang, Yin & Kim, 1996 – China C. tikaderi Majumder & Tikader, 1991 – India C. tixing Yu & Li, 2021 – China C. tongdaoensis Zhang, Yin, Bao & Kim, 1997 – China, Korea C. tongi Yu & Li, 2019 – China C. topakea Barrion & Litsinger, 1995 – Philippines C. torta Forster, 1979 – New Zealand C. tortuosa Zhang & Yin, 1998 – China C. transbaicalica Mikhailov, 1992 – Russia (South Siberia) C. tricuspidata X. Y. Zhang, Li & Z. S. Zhang, 2025 – China C. tridentata Dhali, Roy, Saha & Raychaudhuri, 2016 – India C. trivialis C. L. Koch, 1843 – North America, Europe, Russia (Europe to Far East), China, Japan C. tsurusakii Hayashi, 1987 – Russia (Kurile Is.), Japan C. turongdaliriana (Barrion & Litsinger, 1995) – Philippines C. uenoi Ono, 1986 – Japan C. umbilensis Lessert, 1923 – South Africa C. unanoa Barrion & Litsinger, 1995 – Philippines C. unikarta Barrion & Litsinger, 1995 – Philippines C. uniyali Sarkar, Quasin & Siliwal, 2023 – India (Himalayas) C. upoluensis Marples, 1964 – Samoa C. vachoni Lawrence, 1952 – South Africa C. vacuna L. Koch, 1873 – Papua New Guinea, Australia (Queensland) C. valens Simon, 1897 – South Africa C. venatoria Rainbow & Pulleine, 1920 – Australia (Lord Howe Is.) C. venusae Barrion & Litsinger, 1995 – Philippines C. victoriaensis Barrion & Litsinger, 1995 – Philippines C. vigil Karsch, 1879 – Russia (Sakhalin, Kurile Is.), Korea, China, Japan C. vigillella Strand, 1918 – Japan C. violaceovittata Schenkel, 1936 – China C. wangchengi Yu & Li, 2021 – China C. wolongica Zhu & An, 1999 – China C. wulingensis Yu & Chen, 2017 – China C. xianning Zhong & Yu, 2022 – China C. xiaoci Yu & Li, 2021 – China C. xiaokong Yu & Li, 2021 – China C. yaginumai Hayashi, 1989 – China, Taiwan, Japan C. yangmingensis Hayashi & Yoshida, 1993 – Taiwan C. yanzhii Zhang & Yu, 2020 – China C. yaoi Yu & Li, 2019 – China C. yaroslavi Mikhailov, 2003 – Russia (Far East) C. yasudai Ono, 1991 – Japan C. yejiei Yu & Li, 2021 – China C. yintiaoling X. Y. Zhang, Li & Z. S. Zhang, 2025 – China C. yinyangjian Yu & Li, 2025 – China C. yoshidai Hayashi, 1989 – Japan C. yueya Yu & Li, 2019 – China C. yurii Mikhailov, 2011 – Mongolia C. zacharovi Mikhailov, 1991 – Russia (Far East), Korea C. zandstrai Barrion & Litsinger, 1995 – Philippines C. zhanggureni Yu & Li, 2019 – China C. zhangmuensis Hu & Li, 1987 – China C. zhangyongjingi Li & Blick, 2019 – China C. zhaoi Yu & Li, 2021 – China C. zhengi Yu & Li, 2019 – China C. zhigangi Yu & Li, 2021 – China C. zhui Xu, Yang & Song, 2003 – China C. zilla Dönitz & Strand, 1906 – Japan C. zimmermanni Marples, 1964 – Samoa C. zyuzini Mikhailov, 1995 – Russia (Far East)

See also

• List of Clubionidae species