Enicocephalidae

Enicocephalidae, also called unique-headed bugs and gnat bugs, are a family of around 300 species of the suborder Heteroptera. They are typically 4 mm (0.16 in) long, and found throughout the world. They have an elongated head, constricted in places, hence their head is 'unique'.

Systelloderes maclachlani

Unique-headed bugs Temporal range: Barremian–Present PreꞒ Ꞓ O S D C P T J K Pg N
Phthirocoris magnus female
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Hemiptera
Suborder:	Heteroptera
Superfamily:	Enicocephaloidea
Family:	Enicocephalidae Stål, 1860

They are classified into about 47 genera placed in five subfamilies. The family members can be separated from those of the Aenictopecheidae on the basis of the pronotal division into three lobes (except in the genus Alienates). They also show polymorphism with winged males and wingless or short-winged females.^[1]

Genera

Genera in the family include:

• Alienates Barber, 1953 ^{i c g}
• Boreostolus Wygodzinsky and Stys, 1970 ^{i c g}
• Brevidorsus Kritsky, 1977 ^{i c g}
• Ciucephalus Štys, 1982 ^g
• Disphaerocephalus Cockerell, 1917 ^g
• Enicocephalinus Azar, Fleck, Nel & Solignac, 1999 ^g
• Enicocephalus Westwood, 1838 ^g
• Gourlayocoris ^{c g}
• Henicocephalus ^g
• Henschiella Horvath, 1888 ^g
• Hoplitocoris Jeannel, 1942 ^g
• Hymenocoris Uhler, 1892 ^{i c g}
• Lomagostus Villiers, 1958 ^g
• Monteithostolus Štys, 1981 ^g
• Nesenicocephalus Usinger, 1939 ^{i c g}
• Oncylocotis Stål, 1855 ^g
• Paenicotechys Štys, 1969 ^g
• Paralienates Maldonado-Capriles, Santiago-Blay & Poinar, 1996 ^g
• Phaenicocleus Štys & Banar, 2009 ^g
• Phthirocoris Enderlein, 1904 ^g
• Phthirostenus ^{c g}
• Proboscidopirates Villiers, 1958 ^g
• Pyrenicocephalus Štys, 2010 ^g
• Stenopirates Walker, 1873 ^g
• Systelloderes Blanchard, 1852 ^{i c g b}
• Tornocrusus Kritsky, 1977 ^g
• Xenicocephalus Wygodzinsky & Schmidt, 1991

Data sources: i = ITIS,^[2] c = Catalogue of Life,^[3] g = GBIF,^[4] b = Bugguide.net^[5]

Fossil genera

• †Allocephalocoris Luo et al., 2020 Burmese amber, Myanmar, Cenomanian
• †Disphaerocephalus Cockerell 1917 Burmese amber, Myanmar, Cenomanian
• †Enicocephalinus Azar et al. 1999 Lebanese amber, Barremian^[6] and Escucha Formation, Albian^[7]
• †Paralienates Maldonado Capriles et al. 1996 Dominican amber, Miocene
• †Paenicotechys Cockerell 1917, Burmese amber, Myanmar, Cenomanian
• †Pyrenicocephalus Štys 2010 London Clay, United Kingdom, Ypresian