List of human evolution fossils

The following tables give an overview of notable finds of hominin fossils and remains relating to human evolution, beginning with the formation of the tribe Hominini (the divergence of the human and chimpanzee lineages) in the late Miocene, roughly 7 to 8 million years ago.

As there are thousands of fossils, mostly fragmentary, often consisting of single bones or isolated teeth with complete skulls and skeletons rare, this overview is not complete, but shows some of the most important findings. The fossils are arranged by approximate age as determined by radiometric dating and/or incremental dating and the species name represents current consensus; if there is no clear scientific consensus the other possible classifications are indicated.

The early fossils shown are not considered ancestors to Homo sapiens but are closely related to ancestors and are therefore important to the study of the lineage. After 1.5 million years ago (extinction of Paranthropus), all fossils shown are human (genus Homo). After 11,500 years ago (11.5 ka, beginning of the Holocene), all fossils shown are Homo sapiens (anatomically modern humans), illustrating recent divergence in the formation of modern human sub-populations.

Further information: Chimpanzee–human last common ancestor, Ardipithecus, Sahelanthropus, Homininae, and Hominini

The chimpanzee–human divergence likely took place during around 10 to 7 million years ago.^[1] The list of fossils begins with Graecopithecus, dated some 7.2 million years ago, which may or may not still be ancestral to both the human and the chimpanzee lineage. For the earlier history of the human lineage, see Timeline of human evolution#Hominidae, Hominidae#Phylogeny.

Image	Name	Age (Ma)	Species	Year discovered	Country	Discovered by	Now located at
	El Graeco	7.20[2]	Graecopithecus	1944, 2017	Greece, Bulgaria Site:Pyrgos Vassilissis, Azmaka	Böhme (Tübingen), Spassov (BAS)	Met, Athens; Tübingen, Germany
	TM 266 (Toumai)	7.00–6.00[3]	Sahelanthropus tchadensis	2001	Chad Site:Djurab Desert	Michel Brunet, Alain Beauvilain, Fanone Gongdibe, Mahamat Adoum and Ahounta Djimdoumalbaye	N'Djamena (Chad), BEAC
	BAR 1000'00	6.1–5.7[4]	Orrorin tugenensis	2000	Kenya Site:Lukeino	Martin Pickford, Kiptalam Cheboi, Dominique Gommery, Pierre Mein, Brigitte Senut
	Trachilos footprints	6.05[5]	Made by hominin or hominin-like primate	2002	Greece	Gerard D. Gierliński
	ALA-VP 1/20[6]	5.65±0.150	Ardipithecus kadabba	1997	Ethiopia Site:Middle Awash	Yohannes Haile-Selassie

Further information: Kenyanthropus, Australopithecus, and Paranthropus

Image	Name	Age (Ma)	Species	Year discovered	Country	Discovered by	Now located at
	Ardi	4.40[7]	Ardipithecus ramidus	1994	Ethiopia	Yohannes Haile-Selassie
	Lothagam mandible (KNM-LT 329)[8]	4.60±0.40[9]	Australopithecus anamensis or undetermined Hominidae	1967	Kenya	Arnold Lewis,[10] Bryan Patterson[11][12][13]
	KNM-TH 13150	4.70±0.55[14]	Australopithecus anamensis	1984	Kenya	Kiptalam Cheboi[11]
	KNM-KP 271	4.00[15]	Australopithecus anamensis	1965	Kanapoi, Kenya	Bryan Patterson[11]
	Laetoli Footprints	3.70	Bipedal hominin	1976	Tanzania	Mary Leakey
	LH 4	3.40±0.50	Australopithecus afarensis	1974	Laetoli, Tanzania	Mary Leakey[16]
	KSD-VP-1/1 (Kadanuumuu)	3.58	Australopithecus afarensis	2005	Ethiopia	Yohannes Haile-Selassie
	KT-12/H1 (Abel)	3.50	Australopithecus bahrelghazali	1995	Chad	Mamelbaye Tomalta and Michel Brunet	N'Djamena (Chad), BEAC
	KNM-WT 22944 G-J[17]	3.50	Australopithecus sp.	1990	Kenya	Multinational team	National Museums of Kenya
	KNM-WT 40000 (Flat Faced Man)[18]	3.50-3.20	Kenyanthropus platyops	1999	Lake Turkana (West Lake Turkana), Kenya	Justus Erus and Meave Leakey[19]
	BRT-VP-3/14	3.40±0.10	Australopithecus deyiremeda	2015	Ethiopia	Yohannes Haile-Selassie[20]
	Stw 573 (Little foot)	3.67	Australopithecus prometheus(?)	1994	Sterkfontein, South Africa	Ronald J. Clarke
	DIK-1 (Selam)	3.30	Australopithecus afarensis	2000	Ethiopia	Zeresenay Alemseged
	AL 288-1 (Lucy)	3.20	Australopithecus afarensis	1974	Ethiopia	Tom Gray, Donald Johanson, Yves Coppens and Maurice Taieb	National Museum of Ethiopia
	AL 200-1	3.10±0.10	Australopithecus afarensis	1975	Afar Region, Ethiopia	Donald Johanson Yves Coppens and Maurice Taieb
	AL 129-1	3.10±0.10	Australopithecus afarensis	1973	Afar Region, Ethiopia	Donald Johanson
	AL 444-2[21]	3.00	Australopithecus afarensis	1992	Afar Region, Ethiopia	Yoel Rak
	LD 350-1[22]	2.775±0.025[23]	Homo(?)	2013	Ethiopia	Chalachew Seyoum
	Taung Child 1	3.03–2.61	Australopithecus africanus	1924	Buxton-Norlim Limeworks, South Africa	Raymond Dart	University of the Witwatersrand

Lower Paleolithic: 2.58–0.3 million years old

Further information: Australopithecus, Paranthropus, Homo, and Homo erectus

	Name	Age (Ma)	Species	Date discovered	Country	Discovered by	Now located at
	KNM-WT 17000 (The Black Skull)	2.50	Paranthropus aethiopicus	1985	Kenya	Alan Walker
	BOU-VP-12/130[24]	2.50	Australopithecus garhi	1997	Ethiopia	Yohannes Haile-Selassie
	STS 71[25]	2.61–2.07	Australopithecus africanus	1947	Sterkfontein, South Africa	Robert Broom and John T. Robinson	Ditsong National Museum of Natural History
	STS 52	2.61–2.07	Australopithecus africanus	1947	Sterkfontein, South Africa	Robert Broom	Ditsong National Museum of Natural History
	UR 501 (Uraha jawbone)	2.40±0.10	Homo rudolfensis[26]	1991	Malawi	Tyson Msiska, Timothy Bromage, Friedemann Schrenk
	STS 5 (Mrs. Ples) (STS 14)[27]	2.07[28]	Australopithecus africanus	1947	Sterkfontein, South Africa	Robert Broom	Ditsong National Museum of Natural History
	DNH 134 (Simon)[29]	2.04[29]	Homo erectus	2015	Drimolen Main Quarry, South Africa	Andy Herries' team (excavated by Richard Curtis, Andy Herries, Angeline Leece; reconstructed by Jesse Martin)	University of the Witwatersrand
	DNH 155[30]	2.04–1.95	Paranthropus robustus	2018	Drimolen Main Quarry, South Africa	Andy Herries and Stephanie Baker's team (first found by Samantha Good and excavated by Samantha Good, Angeline Leece, Stephanie Baker and Andy Herries; reconstructed by Jesse Martin)	University of the Witwatersrand
	DNH 152[29](Khethi)	2.04–1.95[29]	Paranthropus robustus	2018	Drimolen Main Quarry, South Africa	Andy Herries and Stephanie Baker's team (first part found by Khethi Nkosi. later parts by Amber Jaeger, Eunice Lalunio; reconstructed by Jesse Martin & Angeline Leece)	University of the Witwatersrand
	DNH 7 (Eurydice)[31]	2.04–1.95[29]	Paranthropus robustus	1994	Drimolen, Drimolen Main Quarry, South Africa	R. Smith and André Keyser	University of the Witwatersrand
	KNM-ER 64060	2.03	Homo habilis	2012	Ileret, Kenya
	KNM-ER 64061	2.02	Homo erectus	2012-2013	Ileret, Kenya
	TM 1517[32]	2.0	Paranthropus robustus	1938	South Africa	Gert Terblanche	Ditsong National Museum of Natural History
	MH1 (Karabo)[33][34]	1.98[35]	Australopithecus sediba	2008	Malapa, South Africa	Matthew Berger and Lee Rogers Berger	University of the Witwatersrand
	KNM-ER 1813	1.90	Homo habilis	1973	Kenya	Kamoya Kimeu
	KNM-ER 1470	1.90	Homo rudolfensis	1972	Kenya	Bernard Ngeneo[36]
	SK 48	2.25–1.80	Paranthropus robustus	1948	Swartkrans, South Africa	Robert Broom	Ditsong National Museum of Natural History
	SK 46[37]	2.25–1.80	Paranthropus robustus	1949	Swartkrans, South Africa	Robert Broom	Ditsong National Museum of Natural History
	SK 847[38]	2.25–1.80	Homo habilis	1949	Swartkrans, South Africa		Ditsong National Museum of Natural History
	OH 24 (Twiggy)[39]	1.80	Homo habilis	1968	Tanzania	Peter Nzube
	OH 8[40]	1.80	Homo habilis	1960	Olduvai, Tanzania
	D2700 (Dmanisi Skull 3)	1.81±0.40[41]	Homo erectus	2001	Dmanisi, Georgia	David Lordkipanidze and Abesalom Vekua
	D3444 (Dmanisi Skull 4)	1.81±0.40	Homo erectus	2003	Dmanisi, Georgia	David Lordkipanidze
	D4500 (Dmanisi Skull 5)	1.81±0.40	Homo erectus	2005 (published in 2013)	Dmanisi, Georgia	David Lordkipanidze
	KNM-ER 62000–62003[42]	1.84±0.60	Homo rudolfensis	2012	Koobi Fora, Kenya	Meave Leakey's team
	KNM-ER 64062	1.84±0.02	Homo erectus	2013	Ileret, Kenya
	OH 5 (Zinj or nutcracker man)	1.75	Paranthropus boisei	1959	Tanzania	Mary Leakey
	OH 7	1.75	Homo habilis	1960	Tanzania	Jonathan Leakey
	StW 53	1.8–1.6[28]	variously A. africanus, H. habilis, H. gautengensis	1976	Sterkfontein, South Africa	A. R. Hughes	University of the Witwatersrand
	KNM-ER 1805	1.74	Homo habilis	1973/4	Kenya	Paul Abell
	Yuanmou Man	1.70 or 0.60–0.50 (disputed)[43]	Homo erectus	1965	China	Fang Qian
	KNM-ER 406	1.70	Paranthropus boisei	1969	Kenya	Richard Leakey
	KNM-ER 732[44]	1.70	Paranthropus boisei	1970	Kenya	Richard Leakey
	KNM-ER 23000[45]	1.70	Paranthropus boisei	1990	Koobi Fora, Kenya	Benson Kyongo
	KNM-WT 17400[46][47]	1.70	Paranthropus boisei	Not known[48]	Lake Turkana (West Lake Turkana) Kenya	unknown[48]	National Museums of Kenya, Nairobi (Kenia)
	KNM-ER 3733	1.63±0.15[49]	Homo ergaster (a.k.a. African Homo erectus)	1975	Kenya
	Lantian Man	1.62±0.03	Homo erectus	1963	Lantian County, China	Woo Ju-Kang
	KNM-WT 15000 (Turkana Boy)	1.60	Homo ergaster (a.k.a. African Homo erectus)	1984	Lake Turkana (West Lake Turkana), Kenya	Kamoya Kimeu	Kenya National Museum
	Peninj Mandible	1.50	Paranthropus boisei	1964	Tanzania	Richard Leakey
	Ileret Footprints	1.50	Homo erectus	2007-2014	Ileret, Kenya
	KNM-ER 992	1.50	Homo ergaster (a.k.a. African Homo erectus)	1971	Kenya	Richard Leakey
	KNM-ER 3883	1.57±0.08	Homo erectus	1976	Kenya	Richard Leakey
	Mojokerto 1 (Mojokerto child)	1.43±0.10	Homo erectus	1936	Indonesia	Andojo, G.H.R. von Koenigswald
	BL02-J54-100[50]	1.40	Similar to H. heidelbergensis	Unknown	Spain	Unknown
	KGA 10-525[51][52]	1.40	Paranthropus boisei	1993	Konso-Gardula, Ethiopia	A. Amzaye
	OH 9 (Chellean Man)[53]	1.40	Homo erectus	1960	Olduvai, Tanzania	Louis Leakey
	Sima del Elephante maxilla[54]	1.40	Homo erectus?	2022	Spain
	ATE9-1[55]	1.20	Homo sp. or Homo erectus?[54]	2008	Spain	Eudald Carbonell	Museo de la Evolución Humana, Burgos (Spain)
	Kocabaş	1.10[56]	Homo erectus[57]	2002	Turkey	M. Cihat Alçiçek
	Daka	1.00	Homo erectus	1997	Ethiopia	Henry Gilbert
	Sangiran 4	1.00	Homo erectus	1939	Indonesia	G.H.R. von Koenigswald
	Sangiran 2	1.15±0.45	Homo erectus	1937	Indonesia	G.H.R. von Koenigswald
	Madam Buya[58]	1.00	Homo erectus	1997	Eritrea	Ernesto Abbate	National Museum of Eritrea
	ATD6-15 and ATD6-69 (Niño de la Gran Dolina 342)	0.900[59]	Homo antecessor or Homo erectus	1994	Spain	Bermúdez & Arsuaga	Museo de la Evolución Humana, Burgos (Spain)
	Trinil 2 Pithecanthropus-1 or Java Man[60]	0.850±0.150	Homo erectus	1891	Indonesia	Eugène Dubois	Naturalis Biodiversity Center, Leiden
	Ternifine 2-3 now Tighennif[61]	0.70	Homo erectus	1954	Algeria	C. Arambourg & B. Hoffstetter
	Sangiran 17[62]	0.70	Homo erectus	1969	Indonesia	S. Sartono
	Peking Man	0.73±0.50[63]	Homo erectus	1921	China	Davidson Black	Lost/stolen
	Nanjing Man	0.60±0.02	Homo erectus	1993	China	Liu Luhong
	Bodo[64]	0.600	Homo heidelbergensis or Homo erectus	1976	Ethiopia	A. Asfaw
	Benjamina[65][66]	0.53	Homo neanderthalensis[67]	2001-2001	Spain	Ana Gracia Téllez
	Mauer 1 (Heidelberg Man)	0.50	Homo heidelbergensis	1907	Germany	Daniel Hartmann	Heidelberg University
	Saldanha man[68]	0.50	Homo rhodesiensis	1953	South Africa
	Boxgrove Man	0.50[69]	Homo heidelbergensis	1994	UK		Natural History Museum
	Arago 21 (Tautavel Man)	0.45	Homo erectus	1971	France	Henry de Lumley
	Ceprano Man[70][71]	0.450±0.050	Homo cepranensis /Homo heidelbergensis	1994	Ceprano, Italy	Italo Biddittu	Servizio di antropologia, Soprintendenza ai beni culturali, Regione Lazio, Italy
	Agamenón[72]	0.43	Homo neanderthalensis[67]	1997	Spain	Paleontological teams	Museo de la Evolución Humana, Burgos (Spain)
	Miguelón	0.40	Homo neanderthalensis[67]	1992	Spain	Bermúdez, Arsuaga & Carbonell	Museo de la Evolución Humana, Burgos (Spain)
	Aroeira 3	0.40	Homo heidelbergensis	2014	Portugal	João Zilhão [de]	Museu Nacional de Arqueologia, Lisbon
	Salé cranium[73][74][75]	0.40-0.20	Homo sapiens?[76]	1971	Morocco	Quarry worker
	Swanscombe Man[77]	0.40	Homo neanderthalensis	1935, 1936, 1955	UK	Alvan T Marston, John J Wymer and Adrian Gibson	Natural History Museum
	Ndutu[78][79]	0.45±.04	Homo neanderthalensis affinities	1973	Tanzania	A.A. Mturi
	Hexian Man[80][81]	0.412±0.025[82]	Homo erectus	1980-1981[83]	Hexian, China
	Gawis cranium	0.350±0.150	Homo erectus/Homo sapiens	2006	Ethiopia	Asahmed Humet
	Steinheim Skull	0.35	Homo heidelbergensis	1933	Germany
	Dinaledi Chamber hominins	0.325±0.090[84]	Homo naledi	2013	South Africa	Rick Hunter and Steven Tucker	University of the Witwatersrand (South Africa)
			Homo heidelbergensis	1949	France		Musée de l'Homme
	BH-1	0.4[85]	Homo heidelbergensis		Mala Balanica, Serbia

Middle Paleolithic: 300,000–50,000 years old

Further information: Middle Stone Age, Southern Dispersal, Archaic humans, and Anatomically modern humans

	Name	Age (ka)	Species	Year discovered	Country	Discovered by	Now located at
	Dragon Man	309–138	Homo longi	1933	China		Hebei GEO University
	Broken Hill 1 (Kabwe 1, Rhodesian Man)	299±25[86]	Homo rhodesiensis (Homo heidelbergensis)	1921	Zambia	Tom Zwiglaar
	Jebel Irhoud 1–5	315±32[87]	Homo sapiens	2017	Morocco	INSAP
	Samu[88]	275±25	Homo heidelbergensis	1964	Hungary	László Vértes
	Dali Man[89]	260±20[90]	Homo daliensis	1978	China	Shuntang Liu
	Jinniushan	260-200 ka[91]	Homo longi Homo daliensis	1984	China[92]	Paleolithic Archeology Student Excavation Team[92]
	Florisbad Skull	259±35	early Homo sapiens or Homo heidelbergensis or Homo helmei	1932	South Africa	T. F. Dreyer, G. Venter
	Galilee Man	250±50	Homo heidelbergensis	1925	Israel	Francis Turville-Petre
	Coupe-Gorge[93]	250