Proctotrupidae

Proctotrupidae is a family of parasitic wasps in the superfamily Proctotrupoidea. There are about 400 species in more than 30 genera in Proctotrupidae, found throughout most of the world.^[2][1]

Proctotrupidae Temporal range: Berriasian–Recent PreꞒ Ꞓ O S D C P T J K Pg N
Codrus picicornis
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Hymenoptera
Suborder:	Apocrita
Infraorder:	Proctotrupomorpha
Superfamily:	Proctotrupoidea
Family:	Proctotrupidae Latreille, 1802
Synonyms
Proctotrypidae[1]

Proctotrupidae are small parasitoid wasps, that primarily parasitise larvae of beetles belonging to the families Carabidae, Staphylinidae, and Elateridae, though a number of species are known to target other hosts, including fungus gnats (Mycetophilidae, Sciaridae) and concealer moths.^[3] Females in many species are wingless and dwell in the soil, where they are capable of detecting beetle larvae in their burrows.^[4] They typically have a body length of 5 to 8 mm, but species may range from 3 to 15 mm.^[2]

Genera

Subfamily Heloriserphinae Engel, Herhold, & Barden, 2022

• Heloriserphus Masner, 1981

Subfamily †Astarteserphinae Engel, Herhold, & Barden, 2022

• †Astarteserphus Engel, Herhold, & Barden, 2022 Lebanese amber Early Cretaceous (Barremian)

Subfamily Austroserphinae Kozlov, 1970

• Austroserphus Dodd, 1933

Subfamily Proctotrupinae Latreille, 1802

tribe Disogmini Kozlov, 1970

• †Cresogmus Rasnitsyn & Kolyada, 2022 Burmese amber, Myanmar, Late Cretaceous (Cenomanian)
• Disogmus Förster, 1856

tribe Cryptoserphini Kozlov, 1970 (=Nothoserphini Kozlov, 1970)

• Brachyserphus Hellèn, 1941
• Cryptocodrus Pschorn-Walcher
• Cryptoserphus Kieffer, 1907
• Hormoserphus Townes & Townes, 1981
• Maaserphus Lin, 1988
• Mischoserphus Townes & Townes, 1981
• Nothoserphus Brues, 1940
• Oxyserphus Masner, 1961
• Phoxoserphus Lin, 1988
• Thomsonina Hellèn, 1941
• Watanabeia Masner, 1958

tribe Proctotrupini Latreille, 1802

• Acanthoserphus Dodd, 1915
• Afroserphus Masner, 1961
• Apoglypha Townes & Townes, 1981
• Austrocodrus Ogloblin, 1960
• Carinaserphus He & Xu, 2007
• Codrus Panzer
• Elgonia Risbec, 1950
• Exallonyx Kieffer, 1904
• Fustiserphus Townes & Townes, 1981
• Glyptoserphus Fan & He, 1993
• Paracodrus Kieffer, 1907
• Parthenocodrus Pschorn-Walcher, 1958
• Phaenoserphus Kieffer, 1908
• Phaneroserphus Pschorn-Walcher, 1958
• Proctotrupes Latreille, 1802
• Pschornia Townes & Townes, 1981
• Serphonostus Townes & Townes, 1981
• Sminthoserphus Townes & Townes, 1981
• Trachyserphus Kolyada, 2017
• Tretoserphus Townes & Townes, 1981

Incertae sedis

• †Dintonia Rasnitsyn & Jarzembowski 1998 Purbeck Group, England, Early Cretaceous (Berriasian)
• †Gurvanotrupes Rasnitsyn, 1986 Gurvan-Eren, Mongolia, Yixian Formation, China, Early Cretaceous (Barremian-Aptian)
• †Pallenites Rasnitsyn & Jarzembowski 1998 Purbeck Group, England, Early Cretaceous (Berriasian)
• †Peverella Rasnitsyn & Jarzembowski 1998 Purbeck Group, England, Early Cretaceous (Berriasian)
• †Protoprocto Sharkey, 1990 Crato Formation, Brazil, Early Cretaceous (Aptian)