Trogiomorpha

Trogiomorpha is one of the three major suborders of barklice, booklice, and parasitic lice in the order Psocodea (formerly Psocoptera),^[2][3] alongside Troctomorpha and Psocomorpha. There are about eight families and more than 430 described species in Trogiomorpha.^[4][5][6] Trogiomorpha is widely agreed to be the earliest diverging of the three suborders, and retains the most primitive characteristics.^[7]

Trogium pulsatorium

Trogiomorpha Temporal range: Barremian–Recent PreꞒ Ꞓ O S D C P T J K Pg N
Dorypteryx domestica
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Psocodea
Suborder:	Trogiomorpha Roesler, 1940
Infraorders[1]
See text

Internal phylogeny

The cladogram below shows the position of Trogiomorpha within Psocodea:^[1]

Psocodea	Troctomorpha Phthiraptera Liposcelididae Pachytroctidae Sphaeropsocidae Amphientometae Psocomorpha Trogiomorpha Atropetae Psyllipsocetae Prionoglaridetae (paraphyletic)

Characteristics

Species of Trogiomorpha possess antennae that are split into 20 or more separate pieces.^[8] The tarsi are segmented into three separate equal pieces, and they have a strong posterior spine. They have been described as having a "mothlike" (brown) appearance in color.^[8]

Classification

Trogiomorpha contains three infraorders and five extant (living) families, as well as three identified extinct families:

• Atropetae
  • †Archaeatropidae Baz & Ortuño, 2000
  • †Empheriidae Baz & Ortuño, 2000
  • Lepidopsocidae Enderlein, 1903 (scaly-winged barklice)
  • Psoquillidae Lienhard & Smithers, 2002 (bird nest barklice)
  • Trogiidae Roesler, 1944 (granary booklice)
• Psyllipsocetae
  • Psyllipsocidae Lienhard & Smithers, 2002 (cave barklice)
• Prionoglaridetae (paraphyletic)
  • Prionoglarididae Azar, Huang & Nel, 2017 (large-winged psocids)
• Unplaced:
  • †Cormopsocidae Yoshizawa & Lienhard, 2020