Boana

Boana is a genus of frogs in the family Hylidae. They are called gladiator frogs, gladiator treefrogs or Wagler Neotropical treefrogs. These frogs live in tropical Central and South America from Nicaragua to Argentina. They also live in the Caribbean, in Trinidad and Tobago.^[1]

Boana
Boana rosenbergi
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Amphibia
Order:	Anura
Family:	Hylidae
Subfamily:	Cophomantinae
Genus:	Boana Gray, 1825
Type species
Rana boans Linnaeus, 1758
Diversity
About 90 species (see text)
Synonyms
List Hysaplesia Boie in Schlegel, 1826 Hypsiboas Wagler, 1830 Auletris Wagler, 1830 Lobipes Fitzinger, 1843 Hypsipsophus Fitzinger, 1843 Phyllobius Fitzinger, 1843 Centrotelma Burmeister, 1856 Hylomedusa Burmeister, 1856 Cinclidium Cope, 1867 Cophomantis Peters, 1870 Cincloscopus Cope, 1871

For a while, scientists thought using this genus was a mistake. They started using it again after taking a long look again at Hylidae. They decided that about 70 species that they had put in the genus Hyla should be in Boana instead. Since then, scientists have written reports about many newly discovered Boana species.^[1][2]

Species

As of April 2020, there are 95 species of frogs in this genus:^[1]

• Boana aguilari (Lehr, Faivovich, and Jungfer, 2010)
• Boana albomarginata (Spix, 1824) – white-edged tree frog
• Boana albonigra (Nieden, 1923) – white-black tree frog
• Boana albopunctata (Spix, 1824) – yellow-spotted tree frog
• Boana alemani (Rivero, 1964) – Cagua tree frog
• Boana alfaroi (Caminer and Ron, 2014)
• Boana almendarizae (Caminer and Ron, 2014)
• Boana appendiculata (Boulenger, 1882)
• Boana atlantica (Caramaschi and Velosa, 1996)
• Boana balzani (Boulenger, 1898) – Yungas tree frog
• Boana bandeirantes (Caramaschi and Cruz, 2013)
• Boana beckeri (Caramaschi and Cruz, 2004)
• Boana bischoffi (Boulenger, 1887) – Bischoff's tree frog
• Boana boans (Linnaeus, 1758) – rusty tree frog
• Boana botumirim (Caramaschi, Cruz, and Nascimento, 2009)
• Boana buriti (Caramaschi and Cruz, 1999)
• Boana caiapo (Pinheiro, Cintra, Valdujo, Silva, Martins, Silva, and Garcia, 2018)
• Boana caingua (Carrizo, 1991) – striped tree frog
• Boana caipora (Antunes, Faivovich, and Haddad, 2008)
• Boana calcarata (Troschel, 1848) – Troschel's tree frog or, convict tree frog
• Boana callipleura (Boulenger, 1902) – Charuplaya tree frog
• Boana cambui (Pinheiro, Pezzuti, Leite, Garcia, Haddad, and Faivovich, 2016)
• Boana cinerascens (Spix, 1824) – Demerara Falls tree frog
• Boana clepsydra (Lutz, 1925) – São Paulo tree frog
• Boana cipoensis (Lutz, 1968)
• Boana cordobae (Barrio, 1965) - Córdoba tree frog
• Boana crepitans (Wied-Neuwied, 1824) – emerald-eyed tree frog
• Boana curupi (Garcia, Faivovich, and Haddad, 2007)
• Boana cymbalum (Bokermann, 1963) – Campo Grande tree frog
• Boana dentei (Bokermann, 1967) – Amapa tree frog
• Boana diabolica (Fouquet et al., 2016)
• Boana ericae (Caramaschi and Cruz, 2000)
• Boana exastis (Caramaschi and Rodrigues, 2003)
• Boana faber (Wied-Neuwied, 1821) – blacksmith tree frog
• Boana fasciata (Günther, 1858) – Gunther's banded tree frog
• Boana freicanecae (Carnaval and Peixoto, 2004)
• Boana geographica (Spix, 1824) – map tree frog
• Boana gladiator (Köhler et al., 2010) - Cusco gladiator tree frog
• Boana goiana (Lutz, 1968)
• Boana guentheri (Boulenger, 1886) – Gunther's Brazilian tree frog
• Boana heilprini (Noble, 1923) – Los Bracitos tree frog
• Boana hobbsi (Cochran and Goin, 1970) – Hobbs' tree frog
• Boana hutchinsi (Pyburn and Hall, 1984) – Hutchins' tree frog
• Boana icamiaba (Pelosoet al., 2018)
• Boana jaguariaivensis (Caramaschi, Cruz, and Segalla, 2010)
• Boana jimenezi (Señaris and Ayarzagüena, 2006)
• Boana joaquini (Lutz, 1968)
• Boana lanciformis (Cope, 1871) – basin tree frog
• Boana latistriata (Caramaschi and Cruz, 2004)
• Boana lemai (Rivero, 1972) – Lema tree frog
• Boana leptolineata (Braun and Braun, 1977) – fine-lined tree frog
• Boana leucocheila (Caramaschi and Niemeyer, 2003)
• Boana lundii (Burmeister, 1856) – Usina tree frog
• Boana maculateralis (Caminer and Ron, 2014)
• Boana marginata (Boulenger, 1887) – Mundo Novo tree frog
• Boana marianitae (Carrizo, 1992) – Salta tree frog
• Boana melanopleura (Boulenger, 1912) – Lower Andes tree frog
• Boana microderma (Pyburn, 1977) – smallskin tree frog
• Boana multifasciata (Günther, 1859) – many-banded tree frog
• Boana nigra (Caminer and Ron, 2020)
• Boana nympha (Faivovich, Moravec, Cisneros-Heredia, and Köhler, 2006)
• Boana ornatissima (Noble, 1923) – ornate tree frog
• Boana palaestes (Duellman, De la Riva, and Wild, 1997)
• Boana paranaiba (Carvalho and Giaretta, 2010)
• Boana pardalis (Spix, 1824) – leopard tree frog
• Boana pellucens (Werner, 1901) – palmar tree frog
• Boana phaeopleura (Caramaschi and Cruz, 2000)
• Boana platanera
• Boana picturata (Boulenger, 1899) – Imbabura tree frog
• Boana poaju (Garcia, Peixoto, and Haddad, 2008)
• Boana polytaenia (Cope, 1870) – Cope's eastern Paraguay tree frog
• Boana pombali (Caramaschi, Pimenta, and Feio, 2004)
• Boana prasina (Burmeister, 1856) – Burmeister's tree frog
• Boana pugnax (Schmidt, 1857) – Chirique-Flusse tree frog
• Boana pulchella (Duméril and Bibron, 1841) – Montevideo tree frog
• Boana pulidoi (Rivero, 1961) – Benitez's tree frog
• Boana punctata (Schneider, 1799) – polka-dot tree frog
• Boana raniceps (Cope, 1862) – Chaco tree frog
• Boana rhythmica (Señaris and Ayarzagüena, 2002)
• Boana riojana (Koslowsky, 1895)
• Boana roraima (Duellman and Hoogmoed, 1992) – Hoogmoed's tree frog
• Boana rosenbergi (Boulenger, 1898) – Rosenberg's tree frog
• Boana rubracyla (Cochran and Goin, 1970) – Valle tree frog
• Boana rufitela (Fouquette, 1961) – Canal Zone tree frog
• Boana secedens (Lutz, 1963) – Barro Branco tree frog
• Boana semiguttata (Lutz, 1925) – speckled tree frog
• Boana semilineata (Spix, 1824)
• Boana sibleszi (Rivero, 1972) – La Escalera tree frog
• Boana steinbachi (Boulenger, 1905)
• Boana stellae (Kwet, 2008)
• Boana stenocephala (Caramaschi and Cruz, 1999)
• Boana tepuiana (Barrio-Amorós and Brewer-Carias, 2008)
• Boana tetete (Caminer and Ron, 2014) - Tetete tree frog
• Boana ventrimaculata (Caminer and Ron, 2020) - Yasuní tree frog
• Boana wavrini (Parker, 1936) – Upper Orinoco tree frog
• Boana xerophylla (Duméril and Bibron, 1841) - emerald-eyed tree frog
  • Boana fuentei (Goin and Goin, 1968) – Fuente's Powakka tree frog, a population of emerald-eyed tree frogs

Hyla palliata and Hypsiboas hypselops are also listed in this article because scientists are not sure about them.^[1]