2009 in paleomammalogy

This paleomammalogy list records new fossil mammal taxa that were described during the year 2009, as well as notes other significant paleomammalogy discoveries and events which occurred during that year.

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Mammals

• A study by J. R. Foster is published estimating the body masses of mammals from the Late Jurassic Morrison Formation by using the ratio of dentary length to body mass of modern marsupials as a reference. Foster concludes that Docodon was the most massive mammal genus of the formation at 141g and Fruitafossor was the least massive at 6g. The average Morrison mammal had a mass of 48.5g. A graph of the body mass distribution of Morrison mammal genera produced a right-skewed curve, meaning that there were more low-mass genera.^[1]
• Fujiwara, S. I. (2009). "Olecranon orientation as an indicator of elbow joint angle in the stance phase, and estimation of forelimb posture in extinct quadruped animals". Journal of Morphology. 270 (9): 1107–1121. doi:10.1002/jmor.10748. PMID 19378290. S2CID 23878624.
• Fujiwara, S. I.; Kuwazuru, O.; Inuzuka, N.; Yochikawa, N. (2009). "Relationship between scapular position and structural strength of rib cage in quadruped animals". Journal of Morphology. 270 (9): 1084–1094. doi:10.1002/jmor.10744. PMID 19378269. S2CID 5574036.
• Mitchell, G.; van Sittert, S.J.; Skinner, J.D. (2009). "Sexual selection is not the origin of long necks in giraffes". Journal of Zoology. 278 (4): 281–286. doi:10.1111/j.1469-7998.2009.00573.x.

Newly named mammals
Name	Status	Authors	Age	Unit	Location	Notes	Images
Anoiapithecus[2]	Valid	Moyà-Solà Alba et al.[CAL 1]	Middle Miocene		Spain
Apatemys pygmaeus[3]	Valid	Beard & Dawson	Early Wasatchian		United States	A member of the family Apatemyidae.
Arcantiodelphys[4]	Valid	Vullo Gheerbrant et al.[CAL 2]	Cenomanian		France
Arvicanthis broekhuisi[5]	Valid	Hordijk De Bruijn	Latest Miocene or earliest Pliocene	Komnina Formation	Greece	A rodent belonging to the family Muridae, a species of Arvicanthis.
Bonisicyon[6]	Valid	Werdelin & Simpson	Late Miocene		Ethiopia  Kenya	A bear dog. The type species is Bonisicyon illacabo.
Choctawius[3]	Valid	Beard & Dawson	Early Wasatchian		United States	A member of Primatomorpha belonging to the family Microsyopidae. The type species is C. foxi; genus also includes "Navajovius" mckennai Szalay (1969).
Colpocherus[3]	Valid	Beard & Dawson	Early Wasatchian		United States	A member of the family Amphilemuridae. The type species is C. mississippiensis.
Corbarimys? nomadus[3]	Valid	Beard & Dawson	Early Wasatchian		United States	A rodent related to members of the genus Paramys.
Corriebaatar[7]	Valid	Rich Vickers-Rich et al.[CAL 3]	Aptian	Wonthaggi Formation	Australia	First Australian multituberculate.
Darwinius[8]	Valid	Franzen Gingerich et al.[CAL 4]	early Geiseltalian	Messel Formation	Germany		Darwinius
Diaceratherium massiliae[9]	Disputed	Ménouret & Guérin	Late Oligocene		France	A rhinoceros. Originally described as a species of Diaceratherium; Tissier, Antoine & Becker (2021) interpreted it as a junior synonym of Ronzotherium romani.[10]
Diacocherus dockeryi[3]	Valid	Beard & Dawson	Early Wasatchian		United States	A member of Erinaceomorpha belonging to the group Litocherinae.
Disallomys[11]	Valid	Korth	Late Oligocene		United States	An aplodontiid rodent, a new genus for the species "Allomys" storeri Tedrow and Korth. Genus also contains two new species: Disallomys robustus and D. intermedius.
Douglassciurus sapphirus[12]	Valid	Korth	Late Oligocene		United States	A sciurine squirrel, a species of Douglassciurus.
Duerotherium[13]	Valid	Cuesta Badiola	Middle Eocene			An anoplotheriine artiodactyl
Ectocion nanabeensis[3]	Valid	Beard & Dawson	Early Wasatchian		United States
Eogale[3]	Valid	Beard & Dawson	Early Wasatchian		United States	A member of the family Miacidae. The type species is E. parydros.
Eritherium[14]	Valid	Gheerbrant	Early Thanetian	Ouled Abdoun basin	Morocco	The oldest, smallest and most primitive elephant relative.
Franimys? actites[3]	Valid	Beard & Dawson	Early Wasatchian		United States	A rodent related to members of the genus Paramys.
Ganlea[15]	Valid	Beard Marivaux et al.[CAL 5]	late Middle Eocene	Pondaung Formation	Myanmar
Haplomylus meridionalis[3]	Valid	Beard & Dawson	Early Wasatchian		United States	A member of the family Hyopsodontidae.
Heliscomys walshi[16]	Valid	Kelly	Middle Eocene	Sespe Formation	United States	A heliscomyid rodent, a species of Heliscomys.
Howellictis[17]	Valid	De Bonis et al.	Late Miocene		Chad	A member of Mustelidae belonging to the subfamily Mellivorinae. The type species is H. valentini.
Kahawamys[18]	Valid	Stevens Holroyd et al.[CAL 6]	Late Oligocene	Nsungwe Formation	Tanzania	A thryonomyoid rodent
Legionarictis[19]	Valid	Tseng Wang Stewart	Miocene (Barstovian)	Temblor Formation	United States	A mustelid. The type species is Legionarictis fortidens.
Maiacetus[20]	Valid	Gingerich ul-Haq et al.[CAL 7]	early Middle Eocene	Habib Rahi Formation	Pakistan		Maiacetus
Maddenia[21]	Valid	Kramarz Bond	Late Oligocene	Sarmiento Formation	Argentina	An astrapothere astrapotherid.
Meiconodon[22]	Valid	Kusuhashi Hu et al.[CAL 8]	Aptian/Albian	Fuxin Formation Shahai Formation	China	A alticonodontine triconodontid
Miacis igniculus[3]	Valid	Beard & Dawson	Early Wasatchian		United States
Microtodon komanensis[5]	Valid	Hordijk De Bruijn	Early Pliocene	Ptolemais Formation	Greece	A rodent.
Mimoperadectes houdei[23]	Valid	Horovitz et al.	Early Wasatchian	Willwood Formation	United States
Mimoperadectes sowasheensis[3]	Valid	Beard & Dawson	Early Wasatchian		United States
Miosengi[24]	Valid	Grossman Holroyd	Early Miocene	Lothidok Formation	Kenya	An elephant shrew. The type species is M. butleri.
Mustela nivalis kudarensis[25]	Valid	Baryshnikov	Middle to late Pleistocene		South Ossetia	A fossil subspecies of the least weasel.
Myomimus igdeliensis[26]	Valid	Suata-Alpaslan	Early Pliocene (Ruscinian)		Turkey	A dormouse, a species of Myomimus.
Mytonolagus ashcrafti[27]	Valid	Fostowicz-Frelik & Tabrum	Eocene		United States	A member of the family Leporidae.
Nalameryx[28]	Valid	Métais Welcomme Ducrocq	Middle Oligocene	Chitarwata Formation	Pakistan	A lophiomerycid ruminant
Naranius americanus[3]	Valid	Beard & Dawson	Early Wasatchian		United States	A member of the family Cimolestidae.
Niglarodon brachyodon[11]	Valid	Korth	Late Oligocene		United States	An aplodontiid rodent, a species of Niglarodon.
Notiolofos[29]	Valid	Bond et al.	Eocene	La Meseta Formation	Antarctica (Seymour Island)	A member of Litopterna belonging to the family Sparnotheriodontidae; a replacement name for Notolophus Bond et al. (2006).[30]
Occitanomys (Rhodomys) vandami[31]	Valid	Suata-Alpaslan	Early Pliocene (Ruscinian)		Turkey	A member of the family Muridae.
Ochotona mediterranensis[32]	Valid	Suata-Alpaslan	Early Pliocene (Ruscinian)		Turkey	A pika.
Oligospermophilus emryi[12]	Valid	Korth	Late Oligocene		United States	A cedromurine squirrel, a species of Oligospermophilus emryi.
Palaeosinopa aestuarium[3]	Valid	Beard & Dawson	Early Wasatchian		United States
Paramys dispar[3]	Valid	Beard & Dawson	Early Wasatchian		United States
Praesinomegaceros venustus[33]	Valid	Vislobokova	Late Miocene (late Turolian)		Russia	A megacerine deer, a species of Praesinomegaceros.
Proconsul meswae[34]	Valid	Harrison Andrews	Early Miocene		Kenya	A species of Proconsul[34] or Ugandapithecus.[35]
Prosciurus clausulus[11]	Valid	Korth	Late Oligocene		United States	An aplodontiid rodent, a species of Prosciurus.
Proterix minimus[36]	Valid	Korth	Late Oligocene		United States	An erinaceid, a species of Proterix.
Pseudomeriones hansi[37]	Valid	Suata-Alpaslan	Early Pliocene (Ruscinian)		Turkey	A gerbil.
Puijila[38]	Valid	Rybczynski Dawson Tedford	Early Miocene		Canada	Extinct genus of pinniped.	Puijila
Sallamys quispea[39]	Valid	Shockey et al.	Deseadan	Moquegua Formation Salla Beds[40]	Bolivia[40]  Peru	A caviomorph rodent related to the group Octodontoidea. Originally described as a species of Sallamys; subsequently transferred to the genus Migraveramus by Pérez et al. (in press).[40]
Simplomys[41]	Valid	García-Paredes in García-Paredes, Peláez-Campomanes & Álvarez-Sierra	Miocene		France  Germany  Portugal  Spain   Switzerland	A dormouse. The type species is "Pseudodryomys" simplicidens De Bruijn (1966); genus also includes "Pseudodryomys" robustus De Bruijn (1967), "Pseudodryomys" aljaphi Hugueney et al. (1978) and "Pseudodryomys" julii Daams (1989), as well as new species S. meulenorum.
Viverriscus[3]	Valid	Beard & Dawson	Early Wasatchian		United States	A member of the family Viverravidae. The type species is V. omnivorus.
Wyonycteris primitivus[3]	Valid	Beard & Dawson	Early Wasatchian		United States	A member of Soricomorpha belonging to the family Nyctitheriidae.

Notes

1. Moyà-Solà, Alba, Almécija, Casanovas-Vilar, Köhler, De Esteban-Trivigno, Robles, Galindo, Fortuny.
2. Vullo, Gheerbrant, de Muizon, Néraudeau.
3. Rich, Vickers-Rich, Flannery, Kear, Cantrill, Komarower, Kool, Pickering, Trusler, Morton, van Klaveren, Fitzgerald.
4. Franzen, Gingerich, Habersetzer, Hurum, von Koenigswald, Smith.
5. Beard, Marivaux, Chaimanee, Jaeger, Marandat, Tafforeau, Soe, Tun, Kyaw.
6. Stevens, Holroyd, Roberts, O'connor, Gottfried.
7. Gingerich, ul-Haq, von Koenigswald, Sanders, Smith, Zalmout.
8. Kusuhashi, Hu, Wang, Hirasawa, Matsuoka.