Permosynidae

Permosynidae is an extinct family of beetles known from the Middle Permian to Early Cretaceous. It is sometimes treated as a form family, because all known members were described from fossils of isolated elytra with punctate striae.^[2][3][4]

Permosynidae Temporal range: 268.0 to 122.46 Ma PreꞒ Ꞓ O S D C P T J K Pg N [1]
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Coleoptera
Suborder:	†Protocoleoptera
Superfamily:	†Permosynoidea
Family:	†Permosynidae Tillyard, 1924
Genera
See text

Taxonomy

After:^[1]

• Alveolacupes Lin, 1983 (also spelled Alveolicupes)
  • Alveolacupes primus Lin, 1983 – Sanqiutang Formation, China, Late Triassic (Rhaetian)
  • Alveolacupes secundes Lin, 1983 – Sanqiutang Formation, China, Late Triassic (Rhaetian)
• Artematopodites Ponomarenko, 1990
  • Artematopodites crispulus Yan in Yan and Zhang, 2010 – Sharteg Formation, Mongolia, Late Jurassic (Tithonian)
  • Artematopodites insculptus (Zhang, 1997) – Sangonghe Formation, China, Early Jurassic (Toarcian); Xishanyao Formation, China, Middle Jurassic (Aalenian/Bajocian)
  • Artematopodites latissimus Ponomarenko, 2014 – Sharteg Formation, Mongolia, Late Jurassic (Tithonian)
  • Artematopodites latus Ponomarenko, 1990 – Byankino Formation, Russia, Late Jurassic (Tithonian)
  • Artematopodites lepidus Yan in Yan and Zhang, 2010 – Sharteg Formation, Mongolia, Late Jurassic (Tithonian)
  • Artematopodites leskoviensis Yan in Yan and Zhang, 2010 – Leskovskaya Formation, Russia, Early Cretaceous (Barremian)
  • Artematopodites longus (Hong, 1983) – Jiulongshan Formation, China, Middle-Late Jurassic (Callovian/Oxfordian)
  • Artematopodites lozowskii Ponomarenko, 2015 – Vokhma Formation, Russia, Late Permian (Changhsingian)
  • Artematopodites major Ponomarenko, 1990 – Turga Formation, Russia, Early Cretaceous (Aptian)
  • Artematopodites maximus Yan in Yan and Zhang, 2010 – Sharteg Formation, Mongolia, Late Jurassic (Tithonian)
  • Artematopodites prolixus (Zhang, 1997) – Badaowan Formation, China, Early Jurassic (Hettangian/Sinemurian)
  • Artematopodites propinquus (Zhang, 1997) – Badaowan Formation, China, Early Jurassic (Hettangian/Sinemurian)
  • Artematopodites shaanbeiensis Hong, 1995 – Huachi-Huanhe Formation, China, Early Cretaceous (Valanginian)
• Bistrisyne Lin, 1986
  • Bistrisyne tenua Lin, 1986 – Sanqiutang Formation, China, Late Triassic (Rhaetian)
• Delpuentesyne Martins-Neto and Gallego, 2007
  • Delpuentesyne menendezi Martins-Neto and Gallego, 2007 – Potrerillos Formation, Argentina, Middle Triassic (Ladinian)
• Diarcuipenna Lin, 1986
  • Diarcuipenna bennettitophila Ponomarenko, 2011 – Lunz Formation, Austria, Late Triassic (Carnian)
  • Diarcuipenna heterosa Lin, 1986 – Zijiachong Formation, China, Late Triassic (Norian)
• Dinoharpalus Handlirsch, 1906
  • Dinoharpalus coptoclavoides Ponomarenko, 2015 – Hassberge Formation, Germany, Late Triassic (Carnian)
  • Dinoharpalus latus Ponomarenko, 2015 – Vokhma Formation, Russia, Late Permian (Changhsingian)
  • Dinoharpalus liasinus (Giebel, 1856) – Cotham Formation, England, Late Triassic (Rhaetian)
  • Dinoharpalus martynovi Ponomarenko, 2018 – Salarevo Formation, Russia, Late Permian (Changhsingian)
  • Dinoharpalus rugosus Ponomarenko, 1985 – Osinovo Formation, Russia, Early-Middle Jurassic (Toarcian/Aalenian)
• Dzeregia Ponomarenko, 1985
  • Dzeregia ampla Ponomarenko, 1990 – Mangutskaya Formation, Russia, Early Cretaceous (Berriasian)
  • Dzeregia byrrhoides Ponomarenko, 1990 – Ukurei Formation, Russia, Late Triassic (Lithonian)
  • Dzeregia crassa Ponomarenko, 1990 – Turga Formation, Russia, Early Cretaceous (Aptian)
  • Dzeregia juxta (Lin, 1986) – Zijiachong Formation, China, Late Triassic (Norian)
  • Dzeregia lata Ponomarenko, 1985 – Makarova Formation, Russia, Early Jurassic (Toarcian)
  • Dzeregia longa Ponomarenko, 1985 – Tersyukskaya Formation, Russia, Middle Jurassic (Bajocian)
  • Dzeregia pilula Ponomarenko, 1990 – Turga Formation, Russia, Early Cretaceous (Aptian)
  • Dzeregia platis Ponomarenko, 2014 – Sharteg Formation, Mongolia, Late Jurassic (Tithonian)
  • Dzeregia striata Ponomarenko, 1985 – Zhargalant Formation, Mongolia, Early Jurassic (Toarcian)
• Hydrobiites Heer, 1865
  • Hydrobiites anglicus Handlirsch, 1906 – Hasfield, England, Late Triassic (Rhaetian)
  • Hydrobiites bellus (Geinitz, 1894) – Dobbertin, Mecklenburg-Vorpommern, Germany, Early Jurassic (Toarcian)
  • Hydrobiites convexus Ponomarenko, 1985 – Tersyukskaya Formation, Russia, Middle Jurassic (Bajocian)
  • Hydrobiites crassus Ponomarenko, 1985 – Cheremkhovskaya Formation, Russia, Early Jurassic (Toarcian)
  • Hydrobiites dobbertinensis Handlirsch, 1939 – Dobbertin, Mecklenburg-Vorpommern, Germany, Early Jurassic (Toarcian)
  • Hydrobiites giebeli Handlirsch, 1906 – Cotham Formation, England, Late Triassic (Rhaetian)
  • Hydrobiites handlirschi Ponomarenko, 2011 – Lunz Formation, Austria, Late Triassic (Carnian)
  • Hydrobiites liasinus (Giebel, 1856) – Hasfield, England, Late Triassic (Rhaetian)
  • Hydrobiites longus Ponomarenko, 1990 – Kutinskaya Formation, Russian, Early Cretaceous (Barremian)
  • Hydrobiites minor Ponomarenko, 2014 – Sharteg Formation, Mongolia, Late Jurassic (Tithonian)
  • Hydrobiites minutissimus Ponomarenko, 1985 – Itat Formation, Russia, Middle Jurassic (Bajocian/Bathonian)
  • Hydrobiites mongolicus Ponomarenko, 2014 – Sharteg Formation, Mongolia, Late Jurassic (Tithonian)
  • Hydrobiites permianus Ponomarenko, 2018 – Salarevo Formation, Russia, Late Permian (Changhsingian)
  • Hydrobiites punctatostriatus Handlirsch, 1939 – Dobbertin, Mecklenburg-Vorpommern, Germany, Early Jurassic (Toarcian)
  • Hydrobiites purbeccensis (Giebel, 1856) – Lulworth Formation, England, Early Cretaceous (Berriasian)
  • Hydrobiites sulcatus Ponomarenko, 1986 – Gurvan-Eren Formation, Mongolia, Early Cretaceous (Aptian)
  • Hydrobiites tillyardi Ponomarenko, 2011 – Vyazniki Formation, Russia, Late Permian (Changhsingian)
  • Hydrobiites veteranus (Heer, 1852) – Insektenmergel Formation, Switzerland, Early Jurassic (Hettangian)
  • Hydrobiites vladimiri Ponomarenko, 2011 – Vyazniki Formation, Russia, Late Permian (Changhsingian)
• Ischichucasyne Martins-Neto and Gallego, 2005
  • Ischichucasyne cladocosta Martins-Neto and Gallego, 2009 – Los Rastros Formation, Argentina, Late Triassic (Carnian)
  • Ischichucasyne santajuanaensis Martins-Neto and Gallego, 2005 – Santa Juana Formation, Chile, Late Triassic (Carnian)
• Palademosyne Rohdendorf, 1961
  • Palademosyne martynovae Rohdendorf, 1961 – Kazankovo-Markinskaya Formation, Russia, Middle Permian (Wordian)
  • Palademosyne natalensis Ponomarenko, 2005 – Estcourt Formation, South Africa, Late Permian (Changhsingian)
• Permocrossos Martynov, 1932
  • Permocrossos elongatus Martynov, 1932 – Iva-Gora Beds Formation, Russia, Middle Permian (Roadian)
• Permosyne Tillyard, 1924
  • Permosyne affinis Tillyard, 1924 – Croudace Bay Formation, Australia, Late Permian (Changhsingian)
  • Permosyne belmontensis Tillyard, 1924 – Croudace Bay Formation, Australia, Late Permian (Changhsingian)
  • Permosyne dentata Ponomarenko, 2003 – Salarevo Formation, Russia, Late Permian (Changhsingian)
  • Permosyne elongata Ponomarenko, 2005 – Estcourt Formation, South Africa, Late Permian (Changhsingian)
  • Permosyne mitchelli Tillyard, 1924 – Croudace Bay Formation, Australia, Late Permian (Changhsingian)
  • Permosyne rasnitsyni Ponomarenko, 2011 – Vyazniki Formation, Russia, Late Permian (Changhsingian)
• Platycrossos Dunstan, 1923
  • Platycrossos caroli Ponomarenko, 2011 – Lunz Formation, Austria, Late Triassic (Carnian)
  • Platycrossos elongatus Ponomarenko, 1985 – Itat Formation, Russia, Middle Jurassic (Bajocian/Bathonian)
  • Platycrossos latus Ponomarenko, 2014 – Sharteg Formation, Mongolia, Late Jurassic (Tithonian)
  • Platycrossos ligulatus Dunstan, 1923 – Blackstone Formation, Australia, Late Triassic (Norian)
  • Platycrossos longus Ponomarenko, 2014 – Sharteg Formation, Mongolia, Late Jurassic (Tithonian)
  • Platycrossos loxonicus Ponomarenko, 2014 – Sharteg Formation, Mongolia, Late Jurassic (Tithonian)
  • Platycrossos mongolicus Ponomarenko, 2014 – Sharteg Formation, Mongolia, Late Jurassic (Tithonian)
  • Platycrossos ovum Ponomarenko, 2014 – Sharteg Formation, Mongolia, Late Jurassic (Tithonian)
  • Platycrossos petalus Ponomarenko, 1985 – Itat Formation, Russia, Middle Jurassic (Bajocian/Bathonian)
  • Platycrossos punctatus Ponomarenko, 1985 – Itat Formation, Russia, Middle Jurassic (Bajocian/Bathonian)
  • Platycrossos subtumidus Dunstan, 1923 – Blackstone Formation, Australia, Late Triassic (Norian)
  • Platycrossos tumidus (Tillyard, 1916) – Blackstone Formation, Australia, Late Triassic (Norian)
• Pseudorhynchophora Handlirsch, 1906
  • Pseudorhynchophora convexa Ponomarenko, 2008 – Nyadeita Formation, Russia, Early Triassic (Olenekian)
  • Pseudorhynchophora olliffi Handlirsch, 1906 – Blackstone Formation, Australia, Late Triassic (Norian)
• Sakmaracoleus Ponomarenko, 2013
  • Sakmaracoleus orenburgensis Ponomarenko, 2013 – Vyazovka Formation, Russia, Middle Permian (Capitanian)
• Tychtocoleus Rohdendorf, 1961
  • Tychtocoleus europaeus (Ponomarenko, 2003) – Salarevo Formation, Russia, Late Permian (Changhsingian)
  • Tychtocoleus neuburgae Rohdendorf, 1961 – Gramoteinskaya Formation, Russia, Late Permian (Wuchiapingian)
  • Tychtocoleus popovi Ponomarenko, 2018 – Salarevo Formation, Russia, Late Permian (Changhsingian)
• Umkomaasia Zeuner, 1960
  • Umkomaasia depressa Zeuner, 1960 – Molteno Formation, South Africa, Late Triassic (Carnian)