List of dinosaur specimens with preserved soft tissue

There have been some discoveries of unusually well-preserved fossil dinosaur specimens which bear remnants of tissues and bodily structures. Organic tissue was previously thought to decay too quickly to enter the fossil record, unlike more mineralised bones and teeth, however, research now suggests the potential for the long-term preservation of original soft tissues over geological time,^[1] leading to the formulation of various hypotheses regarding the underlying mechanisms involved.^[2]

Ornithischians

Basal ornithischians

Nickname	Catalogue Number(s)	Institution(s)	Taxon	Age	Unit	Country(s)	Notes	Images
	INREC K3/109	Institute of Natural Resources, Ecology, and Cryology	Kulindadromeus zabaikalicus	Bathonian	Ukureyskaya Formation	Russia	A basal ornithischian that preserves protofeathers.

Ornithopods

Nickname	Catalogue Number(s)	Institution(s)	Taxon	Age	Unit	Country(s)	Notes	Images
	AMNH 5240	American Museum of Natural History	Corythosaurus casuarius	Campanian	Dinosaur Park Formation	Alberta, Canada
Trachodon mummy	AMNH 5060	American Museum of Natural History	Edmontosaurus annectens	Maastrichtian	Lance Formation	United States		The "Trachodon mummy"
Duck Bill	SMF R 4036	Naturmuseum Senckenberg	Edmontosaurus annectens	Maastrichtian
Dakota	MRF-03	Marmarth Research Foundation	Edmontosaurus annectens	Maastrichtian	Hell Creek Formation	United States	On display at North Dakota Heritage Center & State Museum	Impression of "Dakota"'s skin
			Edmontosaurus regalis
			Gasparinisaura cincosaltensis
			Gryposaurus notabilis
	MOR 548[3][4]	Museum of the Rockies	Hypacrosaurus stebingeri
			Lambeosaurus lambei
			Lambeosaurus magnicristatus
			Maiasaura peeblesorum
Big Paul	LACM 17715	Natural History Museum of Los Angeles County	Magnapaulia laticaudus	Campanian	El Gallo Formation	Mexico		Impression of Big Paul's skin displayed at LACM
			Parasaurolophus walkeri
	AMNH 5220, AMNH 5221, and AMNH 5271	American Museum of Natural History	Saurolophus osborni	Maastrichtian	Horseshoe Canyon Formation	Canada
	Various	Dinosaurium, among others	Saurolophus angustirostris	Maastrichtian	Nemegt Formation	Mongolia, and China	Many specimens are known from the locale called the Dragon's Tomb, which often perseveres soft tissues. This makes it a target for fossil poachers
Leonardo		The Children's Museum of Indianapolis (on ten-year loan from Great Plains Dinosaur Museum and Field Station, 2013-2023)	Brachylophosaurus canadensis	Campanian	Judith River Formation	United States	Well-preserved mummified specimen	Leonardo the Brachylophosaurus
Wilo			Thescelosaurus neglectus	Maastrichtian	Hell Creek Formation	South Dakota, United States	A controversial specimen supposedly with a fossilized four-chambered heart. This is probably a concretion
			Thescelosaurus sp.	Maastrichtian	Hell Creek Formation, Tanis Site		Currently unpublished hind limb with extensive skin preservation present with scales
	NHMUK R2501[5]	Natural History Museum, London	Iguanodontia indent.	Valanginian	Wealden Group	England	Endocast with preserved fossilized brain tissue. Possibly belongs to Barilium dawsoni or Hypselospinus fittoni

Ceratopsians

Nickname	Catalogue Number(s)	Institution(s)	Taxon	Age	Unit	Country(s)	Notes	Images
	AMNH FARB 5427	American Museum of Natural History	Centrosaurus apertus
	TMP 1986.018.0097		Centrosaurus sp.
	UALVP 52613, and CMN 2245	Canadian Museum of Nature	Chasmosaurus belli
	SMF R 4970	Naturmuseum Senckenberg	Psittacosaurus sp.	Aptian	Jehol Biota	China	This specimen displays pigment preservation, bristles as well as a pigmented cloaca.
	AMNH 6418	American Museum of Natural History	Protoceratops andrewsi	Campanian	Djodochta Formation, Bayn Dzak Locality	Mongolia
Lane	Formerly BHI-6273	Houston Museum of Natural Science[6]	Triceratops horridus	Maastrichtian	Lance Formation	United States	Preserves extensive skin impressions, including large "nipple" scales on the sacrum and dorsum.[6]
			Triceratops sp.	Maastrichtian	Hell Creek Formation, Tanis Site	North Dakota, United States	Currently unpublished

Thyreophorans

Nickname	Catalogue Number(s)	Institution(s)	Taxon	Age	Unit	Country(s)	Notes	Images
Suncor Nodosaur	TMP 2011.033. 0001[7] [8] [9] [10] [11] [12] [13]	Royal Tyrrell Museum of Palaeontology	Borealopelta markmitchelli	Albian	Clearwater Formation	Canada	This animal displays pigment preservation, as well as keratinous sheaths and stomach contents.
Victoria	SMA 0018[14]		Hesperosaurus mjosi	Kimmeridgian	Morrison Formation, Howe Quarry	Wyoming, United States
	QM F1801	Queensland Museum	Kunbarrasaurus ieversi	Albian	Allaru Mudstone Formation	Australia
	IVPP-V-12560		Liaoningosaurus paradoxus	Aptian	Yixian Formation	China
Zuul (originally Sherman)	ROM-75860	Royal Ontario Museum	Zuul crurivastator	Campanian	Judith River Formation	United States

Theropods

Basal theropods

Nickname	Catalogue Number(s)	Institution(s)	Taxon	Age	Unit	Country(s)	Notes	Images
	AMNH FR 7224[15]	American Museum of Natural History	Coelophysis bauri	Norian	Ghost Ranch Locality	New Mexico, United States	Stomach contents with the bones of what was once believed to belong to a juvenile Coelophysis individual, showing evidence of cannibalism. However, more recent reviews suggest these to be the bones of the small Crocodylomorph Hesperosuchus agilis	Coelophysis bauri with crocodylomorph remains in their stomach cavity
			Coelophysis bauri				Another possible example of cannibalism with a supposed juvenile leg bone in an adult Coelophysis stomach. However, further examination reveals that the juvenile was actually deposited below another individual and was not in the stomach cavity at all
	SGDS 15-T1		Eubrontes isp.
			Fulicopus lyellii				An impression of a sitting Dilophosaurus-like theropod that potentially shows feathers or buildups in the mud
	SGDS 642-1165		Grallator isp.
	CUE JJ_M01-3[16]		Minisauripus chuanzhuensis		Haman Formation	South Korea

Avialans

Nickname	Catalogue Number(s)	Institution(s)	Taxon	Age	Unit	Country(s)	Notes	Images
Beijing Specimen	BMNHC PH828[17]	Beijing Museum of Natural History	Anchiornis huxleyi	Oxfordian	Tiaojishan Formation	China	An anchiornithid; smaller specimen with preserved feathers covering the legs and arm with fossilized melanosomes indicating a dark grey and black coloration. May not belong to Anchiornis[18]	Anchiornis huxleyi Beijing specimen
	BMNHC PH804[19]	Beijing Museum of Natural History	Anchiornis huxleyi	Oxfordian	Tiaojishan Formation	China	Nearly complete and articulated specimen with preserved feathers on the tail, forelimb, and hindlimb
	IVPP V14378[20]	Institute of Vertebrate Paleontology and Paleoanthropology	Anchiornis huxleyi	Oxfordian	Tiaojishan Formation	China	Holotype specimen with faint preservation of feathers on the back
	LPM-B00169[21][22]	Liaoning Paleontological Museum	Anchiornis huxleyi	Oxfordian	Tiaojishan Formation	China	Nearly complete and articulated specimen with extensive feather preservationon the wings, legs and feet. Along with long head feathers with preserved pigments which indicate a red color	Anchiornis huxleyi Liaoning Specimen
	YTGP-T5199[18]	Yizhou Fossil & Geology Park	Anchiornis huxleyi	Oxfordian	Tiaojishan Formation	China	Nearly complete specimen with preserved feathers on the legs, tail, and some around the body. There are also preserved melanosomes indicating the presence of eumelanin, implying a dark coloration	Referred Anchiornis huxleyi YTGP-T5199 and accompanying diagram
	PKUP V1068[19]	Peking University	Anchiornis huxleyi	Oxfordian	Tiaojishan Formation	China	Nearly complete specimen with some feather impressions around the tail
	STM-0-38, 224, 227, 228[23]	Shandong Tianyu Museum of Nature	Anchiornis huxleyi	Oxfordian	Tiaojishan Formation	China	Preserved plumage and gastric pellets
	STM-0-214[24][25]	Shandong Tianyu Museum of Nature	Anchiornis huxleyi	Oxfordian	Tiaojishan Formation	China	Nearly complete specimen with preserved scant feathers around the wing and legs
	STM-0-179[23]	Shandong Tianyu Museum of Nature	Anchiornis huxleyi	Oxfordian	Tiaojishan Formation	China	Exceptionally preserved specimen with fossilized plumage and gastric pallet in the oesophageal region containing lizard bones	Anchiornis huxleyi specimen STM-0-179
	STM-0-4, 7, 114, 118, 125, 127, 132, 133, 144, 147[21][26][27][28]	Shandong Tianyu Museum of Nature	Anchiornis huxleyi	Oxfordian	Tiaojishan Formation	China	Multiple specimens with preserved feathers on the arms, legs, and around the body. As well as skin impressions showing scales on the feet, ankles, fingers and claws	Anchiornis huxleyi skeletal diagram by Scott Hartman showing multiple feather impressions from multiple specimens
	YFGP T5198[29]	Yizhou Fossil & Geology Park	Aurornis xui	Oxfordian	Tiaojishan Formation	China	An anchiornithid bird with traces of preserved feathers on the tail, chest, and neck. Possibly synonymous with Anchiornis huxleyi[19]
Urfeder	BSP 1869 VIII and 1MB.Av.100 (counterslab)[30][31][32][33] (See Specimens of Archaeopteryx)	Natural History Museum, Berlin	Archaeopteryx lithographica	Tithonian	Solnhofen Limestone	Germany	A single isolated fossil feather with preserved melanosomes indicating a probable black coloration for at least the wings. Has in the past been considered to belong to another avialan, but recent studies reaffirm Archaeopteryx affinties[32][34]	Archaeopteryx lithographica feather imprint, formerly assigned as the holotype
London Specimen	NHMUK 37001[35][36][37](see Specimens of Archaeopteryx)	Natural History Museum, London	Archaeopteryx lithographica	Tithonian	Solnhofen Limestone	Germany	Body missing the head with preserved feathers on the tail and wings	Archaeopteryx lithographica London Specimen
Maxberg Specimen	S5[35][38] (see Specimens of Archaeopteryx)	Maxberg Museum	Archaeopteryx lithographica	Tithonian	Solnhofen Limestone	Germany	A now lost specimen with light feather impressions on the wings and legs	Archaeopteryx lithographica Maxberg Specimen
Munich Specimen	BSP 1999 I 50[35] (see Specimens of Archaeopteryx)	Munich Paleontological Museum	Archaeopteryx lithographica	Tithonian	Solnhofen Limestone	Germany	Nearly complete specimen with preserved feathers on the tail and wings	"Archaeopteryx bavarica" Munich Specimen
Bürgermeister-Müller Specimen (Chicken Wing)	Counterslab[35]	Bürgermeister-Müller Museum (on an unlimited loan)	cf. Archaeopteryx lithographica	Tithonian	Solnhofen Limestone	Germany	Slight feather impressions on the counterslab around the second finger	Archaeopteryx lithographica Chicken Wing
Altmühl Specimen	[39][35]	Naturmuseum Senckenberg	Archaeopteryx lithographica	Tithonian	Solnhofen Limestone	Germany	Nearly complete specimen with preserved feathers on the body and legs	Archaeopteryx lithographica Altmühl Specimen
Chicago Specimen	PA 830[40] (see Specimens of Archaeopteryx)	Field Museum	Archaeopteryx lithographica	Tithonian	Solnhofen Limestone	Germany	Nearly complete specimen with preserved feathers on the wings and tail	Archaeopteryx lithographica Chicago Specimen
Berlin Specimen	MB.Av.101[35][41][42] (see Specimens of Archaeopteryx)	Natural History Museum, Berlin	Archaeopteryx siemensii	Tithonian	Solnhofen Limestone	Germany	Exceptionally preserved specimen with detailed feather preservation around the wings, tail, legs, and body	Archaeopteryx siemensii famous Berlin Specimen
Thermopolis Specimen	WDC CSG 100[43] (see Specimens of Archaeopteryx)	Wyoming Dinosaur Center	Archaeopteryx siemensii	Tithonian	Solnhofen Limestone	Germany	Well preserved specimen with faintly preserved feathers on the right wing and tail	Archaeopteryx siemensii Thermopolis Specimen
	IVPP V14287[44]	Institute of Vertebrate Paleontology and Paleoanthropology	Archaeorhynchus spathula	Aptian	Yixian Formation	China	A nearly complete articulated subadult euornithine bird with preserved plumage around the body and three dozen gastroliths in the stomach
	IVPP V17075/91[45]	Institute of Vertebrate Paleontology and Paleoanthropology	Archaeorhynchus spathula	Aptian	Jiufotang Formation	China	Two specimens with preserved plumage around the body and gastroliths within the stomach	Archaeorhynchus spathula specimen IVPP V17091
	STM7-11[46]	Shandong Tianyu Museum of Nature	Archaeorhynchus spathula	Aptian	Jiufotang Formation	China	Exceptionally preserved specimen with preserved feathers, approximately 100 gastroliths within the stomach region, and impressions of probable lung tissue
	LNTU-WLMP-18[47]	Western Liaoning Museum of Paleontology	Archaeorhynchus spathula	Aptian	Jiufotang Formation	China	A juvenile specimen with preserved feathers on the wings and gastric contents containing propagules, fruits, seeds, and ovules, although no gastroliths	Archaeorhynchus spathula specimen LNTU-WLMP-18
	STM7-145/163[48]	Shandong Tianyu Museum of Nature	Archaeornithura meemannae	Aptian	Huajiying Formation	China	A hongshanornithid bird with extensively preserved feathers on the wings and body	Archaeornithura meemannae holotype specimen and counterslab
	IVPP V25371[49]	Institute of Vertebrate Paleontology and Paleoanthropology	Avimaia schweitzerae	Aptian	Xiagou Formation	China	Lower body of an enantiornithine with some small patches of feathers preserved near the tail	Avimaia schweitzerae holotype with an unlaid egg
	IVPP V17970[50]	Institute of Vertebrate Paleontology and Paleoanthropology	Bellulornis rectusunguis	Aptian	Jiufotang Formation	China	An ornithuromorph bird with faintly preserved feathers on the right wing and tail, as well as gastroliths within the stomach
	LPM B00167[51]	Liaoning Paleontological Museum	Bohaiornis guoi	Aptian	Jiufotang Formation	China	A bohaiornithid enantiornithine with preserved two ribbon-like tail feathers
	IVPP V 17963[52][53]	Institute of Vertebrate Paleontology and Paleoanthropology	Bohaiornis guoi	Aptian	Jiufotang Formation	China	Large stones, interpreted as gastroliths in the stomach cavity. But are more likely rocks deposited on top of the specimen and are not in the stomach at all
	IVPP V26197[54]	Institute of Vertebrate Paleontology and Paleoanthropology	Brevidentavis zhangi	Aptian	Xiagou Formation	China	An ornithuromorph bird with a possible gastric pellet preserved alongside the vertebrae
	IVPP V13266[55]	Institute of Vertebrate Paleontology and Paleoanthropology	Brevirostruavis macrohyoideus	Aptian	Jiufotang Formation	China	An enantiornithine bird with faintly preserved feathers around the wings, body, and tail
	PMoL-B00175[56]	Paleontological Museum of Liaoning	Caihong juji	Oxfordian	Tiaojishan Formation	China	A probable anchiornithid paravian with preserved plumage around the body that possesses pigments that might indicate black and iridescent “rainbow” colors, similar to that of a common grackle	Caihong juji holotype and counterslab specimens
	AMNH 30560/78[57][58][59]	American Museum of Natural History	Calciavis grandei	Ypresian	Green River Formaton	Wyoming, United States	A lithornithid bird with two specimens each with preserved wing feathers, skin impressions of pes scales, and plumage around the body which preserves pigments indicating a glossy, iridescent black, like that of a crow
	IMG 5191	Beijing Museum of Natural History	Cathayornis aberransis	Aptian	Jiufotang Formation	China	Preserved plumage, wing feathers, and bifurcated tail
	FMNH PA730[60]	Field Museum of Natural History	Celericolius acriala	Ypresian	Green River Formation	Wyoming, United States	An Eocene mousebird with preserved tail feathers
	GMV-2129[61]	Geological Museum of China	Changchengornis hengdaoziensis	Aptian	Yixian Formation	China	A confuciusornithid bird with extensively preserved feathers around the body, wings, and a bifurcated tail	Changchengornis hengdaoziensis holotype and counterslab specimens
	AGB5840[62]	Anhui Paleontological Museum	Changzuiornis ahgmi	Aptian	Jiufotang Formation	China	A gansuid ornithuromorph with feather impressions on the tail and wings, with preserved melanosomes indicating a black coloration, and gastroliths within the stomach	Changzuiornis ahgmi holotype specimen
	STM29-11[63]	Shandong Tianyu Museum of Nature	Chiappeavis magnapremaxillo	Aptian	Jiufotang Formation	China	A pengornithid enantiornithine with preserved wing and tail feathers
	STM9-9[64]	Shandong Tianyu Museum of Nature	Chongmingia zhengi	Aptian	Jiufotang Formation	China	A jinguofortisid basal bird with preserved plumage around the body and gastroliths in the stomach	Chongmingia zhengi holotype specimen
	MCCM-LH-1184[65]	Science Museum of Castilla-La Mancha	Concornis lacustris	Barremian	La Huérguina Formation	Spain	An enantiornithine bird with slight feather impressions on the wings and tail	Concornis lacustris holotype cast
	IVPP V11553[66][67]	Institute of Vertebrate Paleontology and Paleoanthropology	Confuciusornis dui	Aptian	Yixian Formation	China	A now lost female specimen with preserved plumage around the body and keratin on the beak
	DNHM D2454[68]	Dalian Natural Museum	Confuciusornis feducciai	Aptian	Jiufotang Formation	China	A well preserved specimen with preserved plumage and bifurcated tail feathers. Probably a synonym of C. sanctus[69]
	IVPP V13168, V13156[70]	Institute of Vertebrate Paleontology and Paleoanthropology	Confuciusornis sanctus	Aptian	Yixian Formation	China	Nearly complete specimens with preserved plumage, wing feathers, and bifurcated tail feathers. Furthermore, skin impressions on the feet	Confuciusornis sanctus specimen IVPP V13156
	IVPP V10920-10925[71]	Institute of Vertebrate Paleontology and Paleoanthropology	cf. Confuciusornis sanctus	Aptian	Yixian Formation	China	Specimen with scant and slight feather impressions
	IVPP V13313[72]	Institute of Vertebrate Paleontology and Paleoanthropology	Confuciusornis sanctus	Aptian	Jiufotang Formation	China	A well preserved specimen with slight feather impressions and scales of Jinanichthys sp. in the throat
	IVPP V13171[73][74]	Institute of Vertebrate Paleontology and Paleoanthropology	Confuciusornis sanctus	Aptian	Jiufotang Formation	China	Nearly complete specimen with preserved skin impressions and feathers with preserved melanosomes, indicating colors of grey, black, brown, and red, similar to a modern zebra finch
	IVPP V12352[75]	Institute of Vertebrate Paleontology and Paleoanthropology	Confuciusornis sanctus	Aptian	Yixian Formation	China	Nearly complete specimen with preserved keratin on the tip of the beak
	IVPP V110304[76]	Institute of Vertebrate Paleontology and Paleoanthropology	Confuciusornis sanctus	Aptian	Yixian Formation	China	Well preserved specimen with preserved wing feathers
	GMV-2130, 2131, 2132, 2133, 2141, 2142, 2146, 2147, 2154[77]	Geological Museum of China	Confuciusornis sanctus	Aptian	Yixian Formation	China	Well preserved plumage around the bodies
	BMNHC PH766[78]	Beijing Museum of Natural History	Confuciusornis sanctus	Aptian	Yixian Formation	China	Well preserved specimen with plumage and skin impressions of scales on the feet
	BMNHC PH925, 931, 986, and 987[79]	Beijing Museum of Natural History	Confuciusornis sanctus	Aptian	Yixian Formation	China	Well preserved plumage around the bodies and bifurcated tail feathers
	DNHM D2859[80]	Dalian Natural Museum	Confuciusornis sanctus	Aptian	Yixian Formation	China	Well preserved specimen with fossilized plumage around the body
	DNHM D1874[81]	Dalian Natural Museum	Confuciusornis sanctus	Aptian	Yixian Formation	China	A probable female specimen with preserved plumage
	DNHM D2508/2151[82][83]	Dalian Natural Museum	Confuciusornis sanctus	Aptian	Yixian Formation	China	Well preserved specimens with indents of plumage and bifurcated tail feathers
	LPM 0228/9[84]	Liaoning Paleontological Museum	Confuciusornis sanctus	Aptian	Yixian Formation	China	Well preserved specimens with indents of plumage and bifurcated tail feathers
	LPM 0233[85]	Liaoning Paleontological Museum	Confuciusornis sanctus	Aptian	Yixian Formation	China	Well preserved specimen with fossilized plumage around the body
	MCFO 0589[82]	CosmoCaixa Barcelona	Confuciusornis sanctus	Aptian	Yixian Formation	China	Well preserved specimen with fossilized plumage around the body
	NIGPAS-139379[86]	Nanjing Institute of Geology and Palaeontology	Confuciusornis sanctus	Aptian	Yixian Formation	China	Two specimens with preserved plumage around the bodies, one with bifurcated tail feathers and the other without, probably representing a male and female
	HGM 41HIII0400/1[87]	Henan Geological Museum	Confuciusornis sanctus	Aptian	Yixian Formation	China	Well preserved specimen with plumage around the body and bifurcated tail feathers
	MGSF315[88]		Confuciusornis sanctus	Aptian	Yixian Formation	China	A well preserved specimen with wing feathers with preserved pigments possibly indicating a lighter white coloration for the wings
	CUGB P1401[89]	China University of Geosciences	Confuciusornis sp.	Aptian	Yixian Formation	China	Preserved feathers with fossilized melanosomes indicating a darker black coloration	Confuciusornis specimen GUGB P1401
	STM13-133/162[67]	Shandong Tianyu Museum of Nature	Confuciusornis sp.	Aptian	Yixian Formation	China	Well preserved specimens with feather impressions and a keratinous rhampotheca
	UFrJ-Dg 031[90][91]	Federal University of Rio de Janeiro	Cratoavis cearensis	Aptian	Crato Formation	Brazil	A small enantiornithine bird with preserved body feathers and bifurcated tail feathers. Furthermore, two specimens of Dastilbe sp. near the digestive track
	IVPP V31106[92]	Institute of Vertebrate Paleontology and Paleoanthropology	Cratonavis zhui	Aptian	Jiufotang Formation	China	A jinguofortisd basal bird with preserved plumage around the body
	IVPP 21711[93][94][95]	Institute of Vertebrate Paleontology and Paleoanthropology	Cruralispennia multidonta	Hauterivian	Huajiying Formation	China	A small enantiornithine bird with preserve plumage present with fossilized melanosomes, indicating a dark brown or black iridescent coloration	Cruralispennia multidonta holotype specimen
	LPM 00039[96]	Liaoning Paleontological Museum	Dapingfangornis sentisorhinus	Aptian	Jiufotang Formation	China	Nearly complete specimen of an enantiornithine with preserved plumage and long bifurcated tail feathers
	USNM 10908[97][98]	National Museum of Natural History	Diatryma? filifera	Ypresian	Green River Formation	Wyoming, United States	Fossilized impressions of what were believed to be feathers belonging to the flightless galloanserae Diatryma, but were in fact Sedge material belonging to the unique species Cyperacites filiferus	Cyperacites filiferus holotype specimen
	IVPP V20284[99]	Institute of Vertebrate Paleontology and Paleoanthropology	Dingavis longimaxilla	Aptian	Yixian Formation	China	A ornithuromorph bird with preserved gastroliths within the stomach
		Museum of Natural Sciences, Brussels	Dynamopterus tuberculata	Lutetian	Messel Formation	Germany	A fossil cuckoo bird with preserved tail feathers and gastroliths in the stomach
	HPG-15-2[100]	Hupoge Amber Museum	Elektorornis chenguangi	Cenomanian	Hukawng Valley	Myanmar	Hindlimb of a small enantiornithine bird found in Burmese amber with preserved plumage and skin
	LH13500[101][102]	Science Museum of Castilla-La Mancha	Eoalulavis hoyasi	Barremian	La Huérguina Formation	Spain	An enantiornithine bird with faintly preserved wing feathers
	IVPP V11977[103][71]	Institute of Vertebrate Paleontology and Paleoanthropology	Eoconfuciusornis zhengi	Hauterivian	Dabeigou Formation	China	An early confuciusornithid bird with preserved plumage around the body and bifurcated tail feathers
	STM7-144[104][105][106]	Shandong Tianyu Museum of Nature	Eoconfuciusornis zhengi	Hauterivian	Dabeigou Formation	China	Preserved skin impressions of scales on the feet, possibly remnants of an ovarian follicle meaning the individual was female, and extensive plumage preservation with fossilized melanosomes indicating a black iridescent coloration
	SMF-ME 1450[107]	Senckenberg Research Institute	Eocoracias brachyptera	Lutetian	Messel Formation	Germany	A stem roller bird with preserved plumage with fossilized melanosomes indicating a non-iridencent blue and black coloration	Eocoracias brachyptera specimen SMF-ME 1450A
	WDC-CGR-109[108]	Wyoming Dinosaur Center	Eocypselus rowei	Ypresian	Green River Formation	Wyoming, United States	An early apodiform with preserved feathers with fossilized melanosomes indicating a black possibly iridescent coloration
			Eoenantiornis buhleri	Aptian	Yixian Formation	China
			Eofringillirostrum boudreauxi
			Eoglaucidium sp.	Lutetian	Messel Formation	Germany
			Eogranivora edentulata
			Eopengornis martini
			Eosinopteryx brevipenna
			Enantiophoenix electrophyla
			Feitianius paradisi
			Fortipesavis prehendens
			Fortunguavis xiaotaizicus
			Gansus zheni
			Gastornis sp.
			Gretcheniao sinensis
			Hassiavis laticauda
	DNHM D2946		Hongshanornis longicresta
			Holbotia ponomarenkoi
			Houornis caudatus
			Ilerdopteryx viai
			Imparavis attenboroughi
	MUSM 1444		Inkayacu paracasensis	Divisaderan-Tinguirirican	Otuma Formation	Peru	A fossil penguin; pigments are preserved on this specimen.
			Iteravis huchzermeyri
			Jeholornis prima
			Jianchangornis microdonta
			Jianianhualong tengi
	IVPP V24194	Institute of Vertebrate Paleontology and Paleoanthropology	Jinguofortis perplexus	Barremian	Dabeigou Formation	China	A pygostylian bird with feather preservation.
			Juehuaornis zhangi
			Junornis houi
			Kaririavis mater
			Longicrusavis houi
	BMNHC PH826		Longipteryx chaoyangensis
			Liaoxiornis delicatus
			Linyiornis amoena
			Longirostravis hani
			Longusunguis kurochkini
			Masillatrogon pumilio
			Mengciusornis dentatus
			Messelornis nearctica
			Messelornis sp.
			Messelirrisor halcyrostris
			Morsoravis sedilis
			Musivavis amabilis
	FMHH PA 778	Field Museum of Natural History	Nahmavis grandei	Eocene	Green River Formation	United States	A bird from the Eocene lagerstätte of the Green River in Wyoming
			Neobohaiornis lamadongensis
			Noguerornis gonzalezi
			Ningyuansaurus wangi
			Oligocolius psittacocephalon
			Ostromia crassipes
			Orienantius ritteri
	MLP 14-I-10-22[109]		Palaeeudyptes gunnari
			Paleoglaux artophoron
			Parahesperornis alexi
			Parapengornis eurycaudatus
			Paraprefica kelleri
			Paraprotopteryx gracilis
			Parargornis messelensis
			Parvavis chuxiongensis
			Pellornis mikkelseni
			Pedopenna daohugouensis
			Piscivorenantiornis insusitatus
			Piscivoravis lii
			Pumiliornis tessellatus
			Praeornis sharovi
			Primobucco mcgrewi
			Cf. Primotrogon sp.
			Primozygodactylus eunjooae
	IVPP V11665		Protopteryx fengningensis
			Salmila robusta
	HGM 41HIII0405		Sapeornis chaoyangensis
	DNHM D3078		Sapeornis chaoyangensis
			Scaniacypselus szarskii
			Schizooura lii
			Septencoracias morsensis
Siky			Serikornis sungei
			Shanweiniao cooperorum
			Shenqiornis mengi
			Shuilingornis angelai
			Similiyanornis brevipectus
			Tianyuornis cheni
			Tynskya eocaena
			Wellnhoferia grandis
			Xiaotingia zhengi
			Xingtianosaurus ganqi
	STM 9-531		Yanornis martini
			Yixianornis grabaui
			Yuanchuavis kompsosoura
			Yuanjiawaornis viriosus
			Zhongornis haoae
	CUGB P1202		Bohaiornithidae indent.
	HPG-15-1[110]		Enantiornithe indent.				A complete juvenile specimen stuck in Burmese amber
	GSGM-07-CM-001		Enantiornithe indent.
	YLSNHM00813[111]	Yingliang Stone Nature History Museum	Enantiornithe indent.				Both wings and hndlimbs with wing feathers of an anantiornithine bird preserved in Burmese amber
	CUGB G20120001		Enantiornithe indent.
	CUGB P1201		Enantiornithe indent.
	BMNHC PH1154/56[112]		Enantiornithe indent.
			Neoaves indent.		Fur Formation
	CUGB G20100053		Ornithurine indent.

Non-avialan maniraptoriformes

Nickname	Catalogue Number(s)	Institution(s)	Taxon	Age	Unit	Country(s)	Notes	Images
	IVPP V24192[113]	Institute of Vertebrate Paleontology and Paleoanthropology	Ambopteryx longibrachium	Oxfordian	Haifanggou Formation	China	A scansoriopterygid with preserved bat-like skin membrane and feathers
	IVPP V11559[114][115]	Institute of Vertebrate Paleontology and Paleoanthropology	Beipiaosaurus inexpectus	Aptian	Yixian Formation	China	a small basal therizinosauroid; a disarticulated specimen of a subadult with preserved plumage around the tail. As well as what has been interpreted as blood cells but are more likely diagenic structures composed of kaolinite clay	Beipiaosaurus inexpectus feather impressions
	STM 31-1[116][117]	Shandong Tianyu Museum of Nature	Beipiaosaurus inexpectus	Aptian	Yixian Formation	China	Specimen composed of a head, neck, and upper half of the torso with preserved protofeathers around the neck. As well as phosphatized skin flakes revealing corneocytes
	BMNHC PH000911[118]	Beijing Museum of Natural History	Beipiaosaurus inexpectus	Aptian	Yixian Formation	China	Upper body with preserved protofeathers around the neck with fossilized melanosomes, indicating a dark brown coloration
	IVPP V12344[119][120]	Institute of Vertebrate Paleontology and Paleoanthropology	Caudipteryx dongi	Aptian	Yixian Formation	China	A caudipterid oviraptorosaur; a nearly complete specimen missing a head with preserved wing feathers and possible skin impressions
	NGMC 97-4-A/9-A[121]	Geological Museum of China	Caudipteryx zoui	Aptian	Yixian Formation	China	Holotype and paratype specimen with preserved feathers on the tails and arms, as well as gastroliths within the stomach	Caudipteryx zoui holotype specimen
	BPM 0001[122][120]	Beipiao Paleontological Museum	Caudipteryx zoui	Aptian	Yixian Formation	China	Nearly complete specimen with preserved plumage and wing feathers
	IVPP V12430[122][123]	Institute of Vertebrate Paleontology and Paleoanthropology	Caudipteryx cf. zoui	Aptian	Yixian Formation	China	Nearly complete specimen with preserved plumage around the body and wing fethers	Caudipteryx sp. specimen IVPP V 12430
	STM4-3[124]	Shandong Tianyu Museum of Nature	Caudipteryx sp.	Aptian	Yixian Formation	China	Nearly complete specimen missing the head with preserved feathers on the arms and around the tail. Molecule remnants are also present. As well as gastroliths within the stomach	Caudipteryx dongi holotype specimen
	HC B016[125]	Natural History Museum of Los Angeles	Changyuraptor yangi	Aptian	Yixian Formation	China	A microraptorine dromaeosaurid; a nearly complete specimen with preserved feathers and plumage around the body, arms, legs, and tail
	UALVP 59400[126]	University of Alberta	Chirostenotes pergracilis	Campanian	Dinosaur Park Formation	Alberta, Canada	A caenagnathid oviraptorosaur with possible feather impressions on the second digit
Big Mama	MPC-D 100/979[127]	Mongolian Paleontological Center	Citipati osmolskae	Campanian	Nemegt Formation	Mongolia	An oviraptorid with faint traces of an α-keratin sheath over the manual unguals
	IMMNH PV00731[128]	Inner Mongolia Museum of Natural History	Daurlong wangi	Aptian	Longjiang Formation	China	Nearly complete basal Dromaeosaurid with preserved plumage on the back and neck, intestinal tracks similar to Scipionyx, and a frog specimen, probably representing Mesophryne near the feet	Daurlong wangi holotype specimen
	MPC-D 100/85[129]	Mongolian Paleontological Center	Duonychus tsogtbaatari	Cenomanian	Bayan Shireh Formation	Mongolia	A therizinosaurid with a trace of a keratinous sheath on the manual ungual
	IVPP V15471[130]	Institute of Vertebrate Paleontology and Paleoanthropology	Epidexipteryx hui	Callovian	Haifanggou Formation	China	A scansoriopterygid with preserved plumage and four long ribbon-like tail feathers	Epidexipteryx hui holotype specimen
	STM4-1 and 22-6[131][132][133]	Shandong Tianyu Museum of Nature	Incisivosaurus sp.	Aptian	Yixian Formation	China	A basal oviraptorosaur; two juvenile specimens formerally assigned to Similicaudipteryx yixianensis with preserved feathers on the tail, hands, and arms. Furthermore, one specimen possibly shows signs of moulting
	41HIII-0308[134]	Henan Geological Museum	Jianchangosaurus yixianensis	Aptian	Yixian Formation	China	A basal therizinosaurian; a nearly complete juvenile specimen with slight feather impressions around the body	Feather impressions Jianchangosaurus yixianensis
	CAGS-IG-04-0801[135][136]	Chinese Academy of Geological Sciences	Jinfengopteryx elegans	Aptian	Huajiying Formation	China	A troodontid with extensive feather preservation present with a feathered tail fan	Jinfengopteryx elegans holotype specimen
	[137][138]		Magnoavipes isp.	Cenomanian	Naturita Formation	Colorado, United States	Possibly ornithomimosaurian ichnogenus; scale scratch marks from the heel region
	IVPP V13352[139][140]	Institute of Vertebrate Paleontology and Paleoanthropology	Microraptor gui	Aptian	Jiufotang Formation	China	Nearly complete specimen with faint feather impressions on the arms, legs, and tip of the tail	Microraptor gui holotype specimen
	IVPP V17972[141][74]	Institute of Vertebrate Paleontology and Paleoanthropology	Microraptor gui	Aptian	Jiufotang Formation	China	Nearly complete specimen with faintly preserved plumage around the body and stomach contents containing the bones of an enantiornithine bird
	IVPP V13320, 51, 477 and TNP00996[139]	Institute of Vertebrate Paleontology and Paleoanthropology and Tianjin Museum of Natural History	Microraptor gui	Aptian	Jiufotang Formation	China	Well preserved specimens with feather impressions
	BMNHC PH881[142][143]	Beijing Museum of Natural History	Microraptor gui	Aptian	Jiufotang Formation	China	Nearly complete specimen with feather impressions with preserved melanosomes, indicating a black iridescent coloration, similar to that of a crow
	BMNHC Ph763	Beijing Museum of Natural History	Microraptor gui	Aptian	Jiufotang Formation	China	Currently unpublished specimen with preserved feather impressions	Microraptor gui specimen BMNHC Ph763
	QM V1002[144]	Beijing Museum of Natural History	Microraptor gui	Aptian	Jiufotang Formation	China	Well preserved specimen with feather impressions around the body and tail and stomach contents containing the remains of fish vertebra
	LVH 0026[145]	Department of Land and Resources of Liaoning Province	Microraptor hanqingi	Aptian	Jiufotang Formation	China	Well preserved specimen with faint feather impressions on the leg
	IVPP V12330[146]	Institute of Vertebrate Paleontology and Paleoanthropology	Microraptor zhaoianus	Aptian	Jiufotang Formation	China	Well preserved specimen with faint feather impressions
	STM5-32[147]	Shandong Tianyu Museum of Nature	Microraptor zhaoianus	Aptian	Jiufotang Formation	China	Nearly complete specimen with stomach contents containing the remains of the lizard Indrasaurus wangi
	STM5-109[26][148]	Shandong Tianyu Museum of Nature	Microraptor sp.	Aptian	Jiufotang Formation	China	Exceptionally complete specimen with wing feathers, propatagium skin, and scale impressions on the pes and soft feet pads	Foot of Microraptor specimen STM 5-109
	STM5-93/172[148][149]	Shandong Tianyu Museum of Nature	Microraptor sp.	Aptian	Jiufotang Formation	China	Exceptionally preserved specimens with wing feathers, propatagium skin
	STM5-4, 5, 9, 75, 142, 150, 221, 6-62, 86[150][148][149]	Shandong Tianyu Museum of Nature	Microraptor sp.	Aptian	Jiufotang Formation	China	Multiple specimens with well preserved wing feather impressions on the arms and legs
	TMP 2009.110.1, 2008.70.1, and 1995.110.1[151][39]	Royal Tyrrell Museum of Palaeontology	Ornithomimus edmontonicus	Campanian	Dinosaur Park Formation	Alberta, Canada	Three specimens, one a juvenile, with slight feather impressions on the body and arms
	UALVP 52531[152]	University of Alberta Laboratory for Vertebrate Paleontology	Ornithomimus sp.	Campanian	Dinosaur Park Formation	Alberta, Canada	Smoot scaleless skin and feather impressions on the leg
	LHC 7777[153][154][155][138]	Science Museum of Castilla-La Mancha	Pelecanimimus polyodon	Barremian	La Huérguina Formation	Spain	A basal ornithomimosaur; smooth scaleless skin on the head, scale impression on the arm, possible occipital crest, and possible lizard-like dewlap or gular pouch similar to modern pelicans, although now prepared over and mostly lost
	NGMC 2125[156]	National Geological Museum of China	Protarchaeopteryx robusta	Aptian	Yixian Formation	China	Basal oviraptorosaur with faint feathers forming a fan on the tail
	UALVP 55700	University of Alberta Laboratory for Vertebrate Paleontology	Saurornitholestes langstoni	Campanian	Dinosaur Park Formation	Alberta, Canada	Nearly complete dromaeosaurid specimen currently still under preparation with traces of a possible keratinous sheath over the sickle claw
	CAGS-02-IG-gausa-1[157][158]	Chinese Academy of Geological Sciences	Scansoriopteryx heilmanni (Epidendrosaurus ninchengensis)	Oxfordian	Tiaojishan Formation	China	A juvenile scansoriopterygid with preserved feather impressions on the arms and tail and skin impressions of scales on the feet
	IGM 100/977[159][160]	Mongolian Institute of Geology	Shuvuuia deserti	Campanian	Djadochta Formation	Mongolia	An alvarezsaurid with faint traces of feathers containing possible β-keratin, although they may be misidentified calcium phosphate
	IVPP V12811[161][73]	Institute of Vertebrate Paleontology and Paleoanthropology	Sinornithosaurus millenii	Aptian	Yixian Formation	China	A microraptorine dromaeosaurid with preserved plumage containing fossilized melanosomes indicating a black and rufous coloration	Sinornithosaurus millenii holotype specimen
Dave	NGMC 91[162][163]	Geological Museum of China	Sinornithosaurus millenii	Aptian	Yixian Formation	China	Exceptionally preserved subadult specimen with plumage around the body	Sinornithosaurus millenii specimen NGMC 91 "Dave"
	MPC-D100/54[164]	Mongolian Paleontological Center	Velociraptor mongoliensis	Campanian	Djadochta Formation	Mongolia	Stomach contents containing azhdarchid pterosaur remains
	DNHM D2933[165][166]	Dalian Natural Museum	Wulong bohaiensis	Aptian	Jiufotang Formation	China	A microraptorine dromaeosaurid with faintly preserved feather impressions around the body, legs, arms, and tail containing fossilized melanosomes indicating a glossy iridescence	Wulong bohaiensis holotype specimen
	STM31-2[167]	Shandong Tianyu Museum of Nature	Yi qi	Oxfordian	Tiaojishan Formation	China	A scansoriopterygid with preserved plumage around the body and skin membrane on the arms forming bat-like wings	Yi qi holotype specimen
	IVPP V12638[168][169]	Institute of Vertebrate Paleontology and Paleoanthropology	Yixianosaurus longimanus	Aptian	Yixian Formation	China	A problematic species of paravian with preserved feathers impressions around the arms
	JPM-0008[170]	Jinzhou Paleontological Museum	Zhenyuanlong suni	Aptian	Yixian Formation	China	A possible microraptorine dromaeosaurid with preserved feathers on the tail, body, and arms	Zhenyuanlong suni holotype specimen
	DIP-V-15103[171]	Dexu Institute of Paleontology	Coelurosauria indent.	Cenomanian	Hukawng Valley	Myanmar	A tail of a juvenile specimen stuck in amber with preserved protofeathers and possibly proteins
	IVPP V13476[172]	Institute of Vertebrate Paleontology and Paleoanthropology	Microraptoria indent.	Aptian	Jiufotang Formation	China	Preserved feathers on the tibiotarsus and the tarsometatarsus
	NMV P160550[173]	Melbourne Museum	Paraves indent.	Aptian	Koonwarra Fossil Bed	Australia	Preserved protofeathers

"Compsognathids"

Nickname	Catalogue Number(s)	Institution(s)	Taxon	Age	Unit	Country(s)	Notes	Images
Compsognathus. corallestris	MNHN CNJ79[174]	National Museum of Natural History, France	Compsognathus longipes	Kimmeridgian	Canjuers Lagerstätte	France	Bumpy structures on the tail interpreted as scales but are more likely to be aberrant bone outgrowths[138]
	IVPP V 14202[175][176]	Institute of Vertebrate Palaeontology & Paleoanthropology	Huadanosaurus sinensis	Aptian	Yixian Formation	China	Potential juvenile specimen with preserved protofeathers with preserved melanosomes indicating a chestnut or light brown coloration in life	Holotype specimen Huadanosaurus sinensis
	CAGS-IG-02-301[177][178]	Chinese Academy of Geological Sciences	Huaxiagnathus orientalis	Aptian	Yixian Formation	China	Gut contents with partially digested unidentified vertebrate bones	Holotype specimen of Huaxiagnathus orientalis
Borsti	JME Sch 200[179][180][181]	Eichstätt Jura Museum	Juravenator starki	Kimmeridgian	Solnhofen Limestone	Germany	Exceptionally preserved juvenile specimen with scaly skin impressions and feathery filaments on the underside of the tail and leg. Possibly a juvenile megalosaurid[182][180]	Holotype specimen of Juravenator starki
	SMNK 2349 PAL[183]	State Museum of Natural History Karlsruhe	Mirischia asymmetrica	Albian	Romualdo Formation	Brazil	Piece of petrified intestine interpreted as a post-cranial air sac similar to modern birds preserved in between and pubis and ischia
	IVPP V12415[175]	Institute of Vertebrate Palaeontology & Paleoanthropology	Sinosauropteryx lingyuanensis	Aptian	Yixian Formation	China	Preserved feathery plumage surrounding the tail	Holotype of Sinosauropteryx lingyuanensis with preserved plumage
	GMV 2123 and NIGP 127586 (Counter slab)[184][185][186]	Geological Museum of China and Nanjing Institute of Geology and Palaeontology	Sinosauropteryx prima	Aptian	Yixian Formation	China	Exceptionally preserved specimen with fossilized protofeathers running along the back. The feathers even preserve melanosomes, indicating a reddish-brown upper side, lighter white underside, and striped tail	Holotype speciman of Sinosauropteryx prima
	NIGP 127587[187][186]	Nanjing Institute of Geology and Palaeontology	Sinosauropteryx prima	Aptian	Yixian Formation	China	A larger specimen with preserved protofeathers running a long the back present with melanosomes. As well as stomach contents containing a specimen of the small shinisaurian lizard Dalinghosaurus sp.	Sinosauropteryx prima holotype specimen (left) compared to the referred specimen NIGP 127587 (right)
	NGMC/GMV 2124[188]	Geological Museum of China	Cf. Sinosauropteryx sp.	Aptian	Yixian Formation	China	Preserved protofeathers and gut contents containing the jaw bones of two individuals of the symmetrodont Zhangheotherium quinquecuspidens and one of the multituberculate Sinobaatar lingyuanensis. Possibly a tyrannosauroid[175]	Specimen GMV 2124 formerly assigned to Sinosauropteryx prima
	JMP-V-05-8-01[189][178]	Jiangxi Provincial Museum	Sinocalliopteryx gigas	Aptian	Yixian Formation	China	Preserves extensive protofeathers on the body and tail. As well as gut contents of an undigested small dromaeosaurid Sinornithosaurus millenii hindlimb with feathers and gastroliths	Holotype specimen of Sinocalliopteryx gigas
	CAGS-IG-T1[178]	Chinese Academy of Geological Sciences	Sinocalliopteryx gigas	Aptian	Yixian Formation	China	Gut contents containing two individuals of the confuciusornithid bird Confuciusornis sanctus and an unidentified ornithischian dinosaur	Abdominal contents of Sinocalliopteryx gigas specimen CAGS-IG-T1
	SMNK PAL 29241	Formally held in the State Museum of Natural History Karlsruhe in Germany but has since been repossessed by the Plácido Cidade Nuvens Paleontology Museum in Brazil[190]	"Ubirajara jubatus"	Aptian	Crato Formation	Brazil	Preserved filaments on its shoulders. The name "Ubirajara" is a nomen nudum since the paper describing it was withdrawn following controversy around the specimen's legality[191]
Little Doggie/Ambrogio/Ciro	SBA-SA 163760[192][193]	Superintendence for the Archaeological Heritage of Salerno	Scipionyx samniticus	Albian	Pietraroja Plattenkalk	Italy	A near-complete juvenile specimen that preserved exceptional phosphatized internal organs, including intestines, tracheal rings, traces of blood tissues, gut contents including lizard remains, a rectum containing fish scales, and even the impression of where the heart would be. Possibly a juvenile carcharodontosaurid[194]	Holotype specimen of Scipionyx samnticus
	BMMS-BK-11[195][180][138]	Bürgermeister-Müller-Museum	Sciurumimus albersdoerferi	Kimmeridgian	Solnhofen Limestone	Germany	Exceptionally preserved juvenile specimen with fossilized protofeather plumage around the tail and body, as well as skin impressions on the underside of the torso and tail. Possibly a juvenile megalosauroid[182]	Holotype specimen of Sciurumimus albersdoeferi

Tyrannosauroids

Nickname	Catalogue Number(s)	Institution(s)	Taxon	Age	Unit	Country(s)	Notes	Images
	TMP 1994.186.0001[196][197]	Royal Tyrrell Museum of Palaeontology	Albertosaurus sarcophagus	Maastrichtian	Horseshoe Canyon Formation	Alberta, Canada	Skin impressions on the underside of the troso
	TMP 99.186.01[197]	Royal Tyrrell Museum of Palaeontology	Albertosaurus sarcophagus	Maastrichtian	Horseshoe Canyon Formation	Alberta, Canada	Skin impressions on the lower torso region
	RTMP 94.186.1[198]	Royal Tyrrell Museum of Palaeontology	Albertosaurus sarcophagus	Maastrichtian	Horseshoe Canyon Formation	Alberta, Canada	Skin impressions
	TMP 2003.45.88[197]	Royal Tyrrell Museum of Palaeontology	Cf. Albertosaurus sarcophagus	Maastrichtian		Alberta, Canada	Disarticulated possible skin impressions from the lower torso region. Possibly Lambeosaurus-like hadrosaurid[138]
	MOR 590[199][200]	Museum of the Rockies	Daspletosaurus horneri	Campanian	Two Medicine Formation	Montana, United States	Ostelogical correlates for cornified pads and large crocodylian-like scales across the entire face. However, the distribution of foramina in the bone seems to indicate that the extra oral tissue doesn't actually resemble Crocodiles much at all[201][202]	Daspletosaurus horneri holotype skull
	TMP 2001.036.0001[196][138]	Royal Tyrrell Museum of Palaeontology	Daspletosaurus torosus	Campanian	Oldman Formation	Alberta, Canada	Skin impressions from an unidentified region of the body
	OTM 200[203]	Old Trail Museum	Daspletosaurus sp.	Campanian	Two Medicine Formation	Montana, United States	Stomach contents containing juvenile hadrosaurids
	IVPP V14243[204]	Institute of Vertebrate Palaeontology & Paleoanthropology	Dilong paradoxus	Aptian	Yixian Formation	China	A basal tyrannosauroid with preserved protofeather impressions near the tail and jaw
	CMN 11593[138][196]	Canadian Museum of Nature	Gorgosaurus libratus	Campanian	Dinosaur Park Formation	Alberta, Canada	Possible small skin impression on the underside of the mid-section of the tail
	NMC 2120[205]	Canadian Museum of History	Gorgosaurus libratus	Campanian	Dinosaur Park Formation	Alberta, Canada	Skin impressions supposedly showing smooth scaleless "naked" skin like that seen some modern flightless birds. Phil Bell and colleagues report that the impressions were covered in plaster and losin the 1980s[196]
	TMP 2009.12.14[206]	Royal Tyrrell Museum of Palaeontology	Gorgosaurus libratus	Campanian	Dinosaur Park Formation	Alberta, Canada	Stomach contents of a juvenile speciman containing the hind limbs of two individuals of the caenagnathid theropod Citipes elegans
	MN 4802-V[138][207][208]	National Museum of Brazil	Santanaraptor placidus	Albian	Romualdo Formation	Brazil	A now potentially destroyed specimen of a basal tyrannosauroid with skin impressions on the hindlimb, present with large scutellate scales similar to some modern birds
	MPD 107/6A[209][210][211]	Central Museum of Dinosaurs of Mongolia	Tarbosaurus bataar	Maastrichtian	Nemegt Formation	Mongolia	Skin impressions on the thoracic region. Probably belongs to a hadrosaurid, possibly Saurolophus angustirostris[138]
	MPD 100F/12[209]	Central Museum of Dinosaurs of Mongolia	Tarbosaurus bataar	Maastrichtian	Nemegt Formation	Mongolia	Skin impressions on the heel and pad of the foot
Tarbosaurus mummy	[212][213][214]		Tarbosaurus bataar	Maastrichtian	Nemegt Formation	Mongolia	A now lost, probably destroyed, specimen with extensive skin preservation on the upper side of the tail and hips
	[215][216][209][217]		Tarbosaurus bataar	Maastrichtian	Nemegt Formation	Mongolia	A now lost articulated skull with accompanying skin impressions complete with a "wattle or bag of skin under the jaws" similar to some modern lizards. Possibly the same specimen as the Tarbosaurus mummy[218]
Wyrex	HMNS 2006.1743.01[196][219]	Houston Museum of Natural Science	Tyrannosaurus rex	Maastrichtian	Hell Creek Formation	Montana, United States	Skin impressions on the underside of the tail, torso, and neck
Bloody Mary	BHI 6437[220][221] (See Specimens of Tyrannosaurus)	North Carolina Museum of Natural Sciences	Tyrannosaurus rex	Maastrichtian	Hell Creek Formation	Montana, United States	Currently unpublished skin impressions on the metatarsal. This specimen may be a juvenile Tyrannosaurus or belong to a unique taxon Nanotyrannus lancensis[222]	Skin impression of Tyrannosaurus rex from the Dueling Dinosaurs specimen
B-rex	MOR 1125[223][224] (See Specimens of Tyrannosaurus)	Museum of the Rockies	Tyrannosaurus rex	Maastrichtian	Hell Creek Formation	Montana, United States	Supposedly preserved red blood cells and proteins within the femur. However this conclusion is controversial and others have supposed it to be permineralized biofilm[225][226]
	AMNH 5027[200][227] (See Specimens of Tyrannosaurus)	American Museum of Natural History	Tyrannosaurus rex	Maastrichtian	Hell Creek Formation	Montana, United States	Possible osteological correlates on the skull indicating cornified pads on the snout and lacrimal rugose ridges	Skull cast of Tyrannosaurus rex speciman AMNH 5027
Dietrich Speciman			Tyrannosaurus rex	Maastrichtian		United States	Currently unpublished possible skin impressions on the neck. Probably not actual skin impressions, possibly a negative impression from a trionychid turtle
	[228]		Cf. Tyrannosaurus rex	Maastrichtian		United States	Unpublished skin impressions from an unknown part of the body. Possibly hadrosaurid?
	ZCDM V5000[229]	Zhucheng Dinosaur Museum and Erlianhaote Dinosaur Museum	Yutyrannus huali	Aptian	Yixian Formation	China	A basal proceratosaurid tyrannosauroid with preserved plumage surrounding the body

Carnosaurs

Nickname	Catalogue Number(s)	Institution(s)	Taxon	Age	Unit	Country(s)	Notes	Images
Big Al II	SMA 0005[138][230]	Aathal Dinosaur Museum	Allosaurus jimmadseni	Kimmerdigian	Morrison Formation, Howe Quarry	Wyoming, United States	Skin impressions on the proximal part of the tail. May belong to a diplodocid sauropod
	WDC DMQ-A 053[138]	Wyoming Dinosaur Center	Allosaurus jimmadseni	Kimmerdigian	Morrison Formation	Wyoming, United States	A juvenile speciman with skin impressions on the pectoral region of the ribs
	[231][232]	The Children's Museum of Indianapolis	Allosaurus sp.	Kimmerdigian	Morrison Formation, Jurassic Mile Locality	Wyoming, United States	Currently unpublished extensive skin impressions
	MNHUK-R-9951[233]	Natural History Museum, London	Baryonyx walkeri	Berremian	Wealden Group	England	Stomach contents revealing the scales of Scheenstia mantelli and a juvenile iguanodontid
	MCCM-LH 6666[234][235]	Science Museum of Castilla-La Mancha	Concavenator corcovatus	Barremian	La Huérguina Formation	Spain	Skin impressions on the underside of the tail, plantar pads on the underside of the feet, and scutate scales on the legs like those seen in some modern birds	Holotype specimen of Concavenator corcovatus
	ML 565-155[138][236]	Lourinhã Museum	Lourinhanosaurus antunesi	Kimmeridgian	Lourinhã Formation	Portugal	An embryonic skeleton with skin impressions present in the shoulder region
	MIWG 6348[237]	Museum of Isle of Wight Geology	Neovenater salerii	Barremian	Wealden Group	England	Osteological correlates on the snout for a complex sensory system

Ceratosaurs

Nickname	Catalogue Number(s)	Institution(s)	Taxon	Age	Unit	Country(s)	Notes	Images
	MCF-PVPF-236[238][239]	Carmen Funes Municipal Museum	Aucasaurus garridoi	Campanian	Anacleto Formation	Argentina	Osteological correlates for attachment points of tail muscles present on the pelvis
	MACN-CH 894[240][241]	Museo Argentino de Ciencias Naturales	Carnotaurus sastrei	Maastrichtian	La Colonia Formation	Argentina	Skin impressions on the underside of the tail, fragments on the left side of the body and skull	Carnotaurus sastrei skin impressions
	UMNH VP 5278[138]	Utah Museum of Natural History	Ceratosaurus dentisulcatus	Kimmeridgian	Morrison Formation, Cleveland-Lloyd Dinosaur Quarry	Utah, United States	Osteoderms from the tail. C. dentisulcatus may be synonymous with C. nascornis
	FMNH PR 2100[242]	Field Museum of Natural History	Majungasaurus crenatissimus	Maatrichtian	Maevarano Formation	Madagascar	Virtually complete skull with ostegical correlates for extra oral tissue	Majungasaurus crenatissimus FMNH PR 2100 skull cast
	MMCH-PV 48[243][244]	Ernesto Bachmann Municipal Museum	Skorpiovenator bustingorryi	Cenomanian	Huincul Formation	Argentina	Possible osteological correlates for muscle attachment points on the pelvis, legs, and on the correlates for scales on the snout and a neurovascular system on the skull
	MMCh‐PV 69[244]	Ernesto Bachmann Municipal Museum	Abelisauridae indent.	Cenomanian	Candeleros Formation	Argentina	Osteological correlates for muscle attachment points on the pubis

Sauropodomorphs

Diplodocids

Nickname	Catalogue Number(s)	Institution(s)	Taxon	Age	Unit	Country(s)	Notes	Images
	MWC 5537[245]	Museum of Western Colorado	cf. Apatosaurus sp.	Kimmeridgian	Morrison Formation, Mygatt-Moore Quarry	Colorado, United States	Skin impressions	(Figure c) Single scale of cf. Apatosaurus sp. (MWC 5537).
	DNM 2873[246]	Dinosaur National Monument	Barosaurus lentus	Kimmeridgian	Morrison Formation, Carnegie Quarry	Utah, United States	Currently unpublished skin impressions
	ML 414[247][248]	Lourinhã Museum	Dinheirosaurus lourinhanensis	Tithonian	Lourinhã Formation	Portugal	Nearly a hundred stones interpreted as gastroliths in the stomach but are more likely pebbles deposited on top of the specimen
Seismosaurus	NMMNH 3690[249][248]	New Mexico Museum of Natural History	Diplodocus hallorum	Kimmeridgian	Morrison Formation	New Mexico, United States	250 stones interpreted as gastroliths in the stomach but are more likely pebbles deposited on top of the specimen
	MDS-2019-028,[250] and CMC VP 10659	Cincinnati Museum Center	Diplodocus sp.	Kimmeridgian	Morrison Formation, Mother's Day Quarry	Montana, United States	Fossilized skin on a young, possibly juvenile individual present on the underbelly, around a limb, and possibly on the neck and head	Skin impressions from a juvenile Diplodocus sp.
	[251][252][253]		cf. Kaatedocus siberi	Kimmeridgian	Morrison Formation, Howe Quarry	Wyoming, United States	Skin impressions present with dermal spines at the end of the tail

Macronarians

Nickname	Catalogue Number(s)	Institution(s)	Taxon	Age	Unit	Country(s)	Notes	Images
	USNM 15560[254]	National Museum of Natural History	Alamosaurus sanjuanensis	Maastrichtian	North Horn Formation	Utah, United States	A saltasaurid titanosaur with dermal osteoderms
	MDE-C3-192[255]	Dinosaur Museum of Espéraza	Ampelosaurus atacis	Maastrichtian	Marnes Rouges Inférieures Formation	France	Titanosaur dermal osteoderms	Ampelosaurus atacis osteoderm
	SMA 0002[256]	Aathal Dinosaur Museum	Camarasaurus sp.	Kimmeridgian	Morrison Formation, Howe Quarry	Wyoming, United States	A camarasaurid with skin impressions on both hind limbs and left fore limb
	DMNH 39045[257][248]	Denver Museum of Natural History	Cedarosaurus weiskopfae	Valanginian	Cedar Mountain Formation, Yellow Cat Member	Utah, United States	A brachiosaurid with 115 stones of varying sizes interpreted as gastroliths preserved in the stomach cavity, but are more likely stones were deposited on top of the specimen
Judy	AODF 0888[258]	Australian Age of Dinosaurs	Diamantinasaurus matildae	Cenomanian	Winton Formation	Australia	A diamantinasaur with stomach contents of conifers, seed ferns, leaves, and angiosperms
	MPM-PV 1156[259][260]	Padre Molina Museum	Dreadnoughtus schrani	Maastrichtian	Cerro Fortaleza Formation	Argentina	A titanosaur specimen with preserved fossilized collegian and internal proteins as well as possible osteological correlates for soft tissue on the neural spine
	NHMUK R1868[261]	Natural History Museum, London	Haestasaurus becklesii	Tithonian	Wealden Group	England	A basal macronarian with skin impressions on the limb	Skin impressions from Haestasaurus becklesii
	FGGUB R.1410[262]	University of Tübingen	Magyarosaurus dacus	Maastrichtian	Sânpetru Formation	Romania	A titanosaur with dermal osteoderms
	PVL 4017−92, 113, 129, 208, PVLPh 011–1-3[263]	National University of Tucumán	Saltasaurus loricatus	Maastrichtian	Lecho Formation	Argentina	A saltasaurid titanosaur with dermal osteoderms of varying sizes across the torso	Saltasaurus loricatus osteoderm
	MEF-PV 1125/1[264]	Museum of Paleontology Egidio Feruglio	Tehuelchesaurus benitezii	Kimmeridgian	Cañadón Calcáreo Formation	Argentina	A camarasaurid with skin impressions	Skin impressions from Tehuelchesaurus benitezii
	FMNH PR 2021-52 and UA 9875[265]	Field Museum of Natural History and University of Antananarivo	"Titanosaurus madagascariensis"	Maastrichtian	Maevarano Formation	Madagascar	A titanosaur with dermal osteoderms
	FMNH PR 2342 and UA 9331[266]	Field Museum of Natural History and University of Antananarivo	Rapetosaurus krausei	Maastrichtian	Maevarano Formation	Madagascar	A titanosaur with dermal osteoderms from an adult and juvenile specimen
	[267]		Brontopodus cf., pentadactylus	Aalenian	Saltwick Formation	England	Manus and pes skin impressions
	MQE.2023.038[268]	Mongolia Quest Expedition	Brontopodus isp.	Maastrichtian	Nemegt Formation	Mongolia	Forelimb manus skin impressions
	[269][270]		cf. Brontopodus isp.	Albian	Haman Formation	South Korea	Hindlimb skin impressions from footprints
	PVPH-130[271]		Cf. Megaloolithidae	Campanian	Anacleto Formation	Argentina	Disarticulated embryonic skeleton containing skin impressions probably from the backside of the animal	A) Large patch (PVH- 126). B) Detail of rosette structure of PVPH-130.
	AMNH 1959[255]	American Museum of Natural History	Titanosauria indent.	Maastrichtian	Lameta Formation	India	Osteoderms
	MPMA 08-0058-11[272]	Monte Alto Museum of Palaeontology	Titanosauria indent.	Campanian	Adamantina Formation	Brazil	Osteoderms
	HUE-00590-00913[273]	Castilla-la Mancha Science Museum	Titanosauria indent.	Campanian	Lo Hueco Site	Spain	Osteoderms
	MMCERCS-1340[274]		Titanosauria indent.	Maastrichtian	Tremp Formation	Spain	Skin impression from a footprint

See also

• Dinosaur coloration
• Feathered dinosaur
• List of non-avian dinosaur species preserved with evidence of feathers
• Dinosaur mummy
• List of fossils with consumulites

Sources

• Chiappe, Luis M.; Meng, Qingjin (2016). Birds of Stone: Chinese Avian Fossils from the Age of Dinosaurs. Baltimore: Johns Hopkins University Press. ISBN 978-0-297-84156-2.

• Sanders, F.; Manley, K.; Carpenter, K. (2001). "Gastroliths from the Lower Cretaceous sauropod Cedarosaurus weiskopfae". In Tanke, Darren; Carpenter, Ken (eds.). Mesozoic Vertebrate Life: New Research Inspired by the Paleontology of Philip J. Currie. Indiana University Press. pp. 166–180. ISBN 0-253-33907-3